Asteraceae


Anthemis cretica
subsp. cassia
Anthemis L.
Anthemis cretica L. subsp. cassia (BOISS.) GRIERSON
Ömür: Çok yıllık
Yapı: ot
Hayat formu:
İlk çiçeklenme zamanı: 6
Son çiçeklenme zamanı: 8
Habitat: kireçtaşı çağıllık, Juniperus veya Cedrus arası
Minimum yükseklik: 750
Maksimum yükseklik: 2200
Endemik: -
Element: ?
Türkiye dağılımı: Karasal ve G. Anadolu
Genel dağılımı: ?
Bulunduğu kareler:A1 A2 A3 A4 A5 A6 A7 A8 A9 B1 B2 B3 B4 B5 B6 B7 B8 B9 C1 C2 C3 C4 C5 C6 C7
 
A. cretica L., Sp. Pl. 895 (1753) non (L.) Nyman (1854-55). Syn: A. montana L., Sp. PL ed. 2:1261 (1763), nom. illegit. A. orientalis auct. non (L.) Degen, Degen It. Turc. (Exsicc, 1890) non Anacyclus orientalis L.
  Shortly rhizomatous perennial herb, sometimes woody at base, subglabrous or ± densely greyish- or white-pubescent. Stems decumbent, ascending or erect, 10-30(-40) cm, simple or branching. Leaves pinnatisect, up to 5 cm, primary segments usually 3-paired, simple or more often divided into 3-5(-9) oblanceolate ± obtuse secondary segments. Capitula radiate or discoid. Involucregenerally 0.75-1.5 cm broad; phyllaries ovate or lanceolate, ± acute, margins ± scarious, colourless, pale or dark brown. Receptacle often becoming somewhat conical at maturity; paleae lanceolate, carinate. Ray flowers when present white, usually fertile; ligules elliptic, 0.5-1.25 cm. Disc flowers yellow, corolla-tube usually becoming inflated at maturity. Achenes obpyramidal, ± squarish in section, 1.5-3.5 mm, smooth, sulcate or especially the outer subtuberculate. Pappus auriculate or coroniform, ± absent or up to 1 mm, scarious. Type: Hb. Cliff. 415/2!
This is a highly polymorphic species in which a number of different taxa are recognizable and have been variously treated at specific or infraspecific levels by previous authors. The following is a tentative classification but, such is the degree of intergradation, no satisfactory scheme seems possible without some biosystematic study; numerous intermediates occur between the subspecies recognized here; radiate and discoid variants are treated informally and epithets have not been given to them as it has seemed difficult to avoid confusion with those applied, perhaps slightly differently, in Europe. The external distribution has been curtailed due to lack of information. The illegitimacy of the name A. montana L. (previously applied to this widespread species complex) is discussed by Grierson in Notes R.B.G. Edinb. 33: 212 (1974).
1. Leaf segments linear-lanceolate, c. 0.5 mm broad, acute or acuminate (some-times broader but then confined to islands); radiate and discoid plants e. subsp. leucanthemoides
1. Leaf segments oblanceolate or obovate, usually more than 0.5 mm broad, obtuse or subacute
  2. Lower or median leaves usually more than 2 cm
    3. Phyllaries with dark brown or blackish margins; plants radiate and discoid, or radiate only
      4. Taller plants (20-)30-40 cm tall; radiate and discoid plants
        5. Leaf segments 1 .5 mm or less broad b. subsp. albida
        5. Leaf segments (l-)2-3.5 mm broad g. subsp. cassia
      4. Smaller plants 15(-20) cm tall (or less from higher alpine habitats); radiate plants only
          6. Leaves usually glabrous (Ulu Da.) j. subsp. carpatica
          6. Leaves ± densely whitish pubescent (N.E. Anatolia) k. subsp. iberica
    3. Phyllaries with whitish or pale brown margins; plants radiate and discoid
             7. Primary leaf segments divided into 7 or more segments h. subsp. absinthifolia
             7. Primary leaf segments divided into 3 or 5 segments
                8. Stems elongate, 30-45 cm, much branched i. subsp. candicans
                8. Stems 10-20(-40) cm, simple or branched only near base
                   9. Plants usually with several leafy sterile shoots; leaf segments ± contiguous a. subsp. pontica
                   9. Plants without or with few sterile shoots; lear sentents ± distant f. subsp. umbilicata
  2. Lower and median leaves usually less than 2 cm
                      10. Phyllaries with whitish or pale brown margins; radiate or discoid plants
                          11. Indumentum dense, leaves often greyish or white; marginal achenes usually subtuberculate d. subsp. anatolica
                          11. Indumentum ± sparse, leaves green; marginal achenes smooth c. subsp. tenuiloba
                      10. Phyllaries with dark brown or blackish margins; radiate plants only l. subsp. argaea
g. subsp. cassia (Boiss.) Grierson in Notes R.B.G. Edinb. 33:428 (1975). Syn: A. cassia Boiss., Diagn. ser. 1(11): 10 (1849)! A. blancheana Boiss., Fl. Or. 3:292 (1875)! quoad plantas Anatolicas.
Stems ascending, 15-30 cm. Leaves 3-7 cm, ± greyish-white pubescent, pinnati-sect segments simple or ternately divided into lobes up to 12 x 1-2.5 mm. Capitula radiate (in type) or discoid. Involucre 1-1.5 cm broad; phyllaries with narrow pale or dark brown margins. Achenes 2.5 mm, ribbed; auricle c. 0.5 mm. Fl. 6-8. Limestone screes and among Juniper or Cedrus, 750-2200 m.
  Type: [Turkey C5 Hatay] in regione superiori Cassii (Akra Da.), vi 1846, Boissier (iso. K!). Radiate variants.
  Inner & S. Anatolia. A7 Giresun: 9-11 km N. of Şebinkarahisar, 1300 m, Hub.-Mor. 13291! B5 Kayseri: 24 km S. of Pinarbaşi, 1720-1750 m, Hub.-Mor. 12391 (approaching subsp. albida)! B7 Erzincan: Atma (Atmak), Sint. 1890:2205! C3 Isparta: E. shore of Eğridir G., 1060 m, Hub.-Mor. 8692! C4 Içel: 32 km E. of Gülnar, 860 m, Hub.-Mor. 9926! Konya: Bozkir, Tufan De., 1750 m, Çetik 224 (slender form with narrow leaf segments)! C5 Adana: E. of Gülek Boğazi, 2000 m, Sorger 62-71-29! C6 Hatay: 5 km W. of Kislak, 750 m, Watson et al. 838! Discoid variants.
  E. & S. Anatolia. B7 Tunceli: 46 km N. of Tunceli, 1100 m, Hub.-Mor. 15798 (intermediate with subsp. ponticd)! C6 Hatay: N.E. of Latik, 1050 m, Hub.-Mor. 12387! Maraş: pass between Armut Da. and Keş Da., 19 km S. of Göksun, 1650 m, Hub.-Mor. 12385!
  According to Boissier, A. cassia has pale brown-margined phyllaries, whereas those of A. blancheana are dark. There appears to be a complete range of variation between the two states. Some of the syntypes of A. blancheana collected in N. Syria by Blanche have much narrower segments than those from S. Anatolia.