Asteraceae


Anthemis cretica
subsp. umbilicata


Anthemis cretica
subsp. umbilicata
Anthemis L.
Anthemis cretica L. subsp. umbilicata (BOISS. ET HUET) GRIERSON
Ömür: Çok yıllık
Yapı: ot
Hayat formu:
İlk çiçeklenme zamanı: 5
Son çiçeklenme zamanı: 7
Habitat: step, kayalık yamaç
Minimum yükseklik: 180
Maksimum yükseklik: 2000
Endemik: -
Element: ?
Türkiye dağılımı: K. ve Karasal Anadolu,
Genel dağılımı: ?
Bulunduğu kareler:A1 A2 A3 A4 A5 A6 A7 A8 A9 B1 B2 B3 B4 B5 B6 B7 B8 B9 C1 C2 C3 C4 C5 C6 C7
 
A. cretica L., Sp. Pl. 895 (1753) non (L.) Nyman (1854-55). Syn: A. montana L., Sp. PL ed. 2:1261 (1763), nom. illegit. A. orientalis auct. non (L.) Degen, Degen It. Turc. (Exsicc, 1890) non Anacyclus orientalis L.
  Shortly rhizomatous perennial herb, sometimes woody at base, subglabrous or ± densely greyish- or white-pubescent. Stems decumbent, ascending or erect, 10-30(-40) cm, simple or branching. Leaves pinnatisect, up to 5 cm, primary segments usually 3-paired, simple or more often divided into 3-5(-9) oblanceolate ± obtuse secondary segments. Capitula radiate or discoid. Involucregenerally 0.75-1.5 cm broad; phyllaries ovate or lanceolate, ± acute, margins ± scarious, colourless, pale or dark brown. Receptacle often becoming somewhat conical at maturity; paleae lanceolate, carinate. Ray flowers when present white, usually fertile; ligules elliptic, 0.5-1.25 cm. Disc flowers yellow, corolla-tube usually becoming inflated at maturity. Achenes obpyramidal, ± squarish in section, 1.5-3.5 mm, smooth, sulcate or especially the outer subtuberculate. Pappus auriculate or coroniform, ± absent or up to 1 mm, scarious. Type: Hb. Cliff. 415/2!
This is a highly polymorphic species in which a number of different taxa are recognizable and have been variously treated at specific or infraspecific levels by previous authors. The following is a tentative classification but, such is the degree of intergradation, no satisfactory scheme seems possible without some biosystematic study; numerous intermediates occur between the subspecies recognized here; radiate and discoid variants are treated informally and epithets have not been given to them as it has seemed difficult to avoid confusion with those applied, perhaps slightly differently, in Europe. The external distribution has been curtailed due to lack of information. The illegitimacy of the name A. montana L. (previously applied to this widespread species complex) is discussed by Grierson in Notes R.B.G. Edinb. 33: 212 (1974).
1. Leaf segments linear-lanceolate, c. 0.5 mm broad, acute or acuminate (some-times broader but then confined to islands); radiate and discoid plants e. subsp. leucanthemoides
1. Leaf segments oblanceolate or obovate, usually more than 0.5 mm broad, obtuse or subacute
  2. Lower or median leaves usually more than 2 cm
    3. Phyllaries with dark brown or blackish margins; plants radiate and discoid, or radiate only
       4. Taller plants (20-)30-40 cm tall; radiate and discoid plants
          5. Leaf segments 1 .5 mm or less broad b. subsp. albida
          5. Leaf segments (l-)2-3.5 mm broad g. subsp. cassia
       4. Smaller plants 15(-20) cm tall (or less from higher alpine habitats); radiate plants only
             6. Leaves usually glabrous (Ulu Da.) j. subsp. carpatica
             6. Leaves ± densely whitish pubescent (N.E. Anatolia) k. subsp. iberica
     3. Phyllaries with whitish or pale brown margins; plants radiate and discoid
                 7. Primary leaf segments divided into 7 or more segments h. subsp. absinthifolia
                 7. Primary leaf segments divided into 3 or 5 segments
                   8. Stems elongate, 30-45 cm, much branched i. subsp. candicans
                   8. Stems 10-20(-40) cm, simple or branched only near base
                       9. Plants usually with several leafy sterile shoots; leaf segments ± contiguous a. subsp. pontica
                       9. Plants without or with few sterile shoots; lear sentents ± distant f. subsp. umbilicata
  2. Lower and median leaves usually less than 2 cm
                          10. Phyllaries with whitish or pale brown margins; radiate or discoid plants
                               11. Indumentum dense, leaves often greyish or white; marginal achenes usually subtuberculate d. subsp. anatolica
                               11. Indumentum ± sparse, leaves green; marginal achenes smooth c. subsp. tenuiloba
                          10. Phyllaries with dark brown or blackish margins; radiate plants only l. subsp. argaea
f. subsp. umbilicata (Boiss. & Huet) Grierson in Notes R.B.G. Edinb. 33:428 (1975). Syn: A. umbilicata Boiss. & Huet in Boiss., Diagn. ser. 2(3):24 (1856)! Stems 18-25(-40) cm, ± erect, usually with few leafy sterile shoots at base. Leaves 3-4 cm, segments rather distant, 0.7-1 cm, ternate from base. Capitula radiate (in type) or discoid. Involucre 0.75-1 cm broad; phyllaries pale-margined. Achenes 1.5-2 mm, ± sulcate; corona c. 0.25 mm. Fl. 5-7. Steppe and on rocky slopes, 180-2000 m.
  Syntypes: [N.E. Turkey] in valle Kassukla, Huet du Pavilion (G! iso. K!); [A8 Gümüşane] Baiburt, Bourgeau 384 (G!). Radiate variants.
  N. & Inner Anatolia. A3 Zonguldak: Devrek to Dirgine, 180 m, D. 37730! A4 Ankara: Hacikadin De., nr Keçiören, D. 18735! A7 Gümüşane: Tekke (Teke), Sint. 1894:7551! A9 Erzurum: Horasan to Karaurgan, 2000 m, D. 30725! B4 Ankara: 10 km S. of Ankara, M.Zohary & Orshan 2812! B7 Elaziğ: 2 km S. of Maden, 1200 m, lt. Leyd. 1959:1540! Discoid variants. C. Anatolia. A4 Ankara: Kizilcahamam, 1300 m, Sorger 70-5-17! Çankiri:
37 km S. of Çankiri, 600 m, Hub.-Mor. 17296! A5 Çorum: 8 km S. of Iskilip, 800 m, Coode & Jones 1803!
  The lack of sterile basal shoots led Boissier to publish this taxon as a biennial. However, such shoots are present on other specimens which are obviously peren-nial, and possibly the appearance of these shoots may depend on environmental conditions.