Apiaceae

Heracleum pastinacifolium subsp. incanum

Heracleum L. Heracleum pastinacifolium C. KOCH subsp. incanum (BOISS. ET HUET) DAVIS Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 7 Son çiçeklenme zamanı: 9 Habitat: kayalık yamaçlar, sel yatakları, çağıllık, doruklar Minimum yükseklik: 1800 Maksimum yükseklik: 3500 Endemik: endemik Element: ? Türkiye dağılımı: K. ve D. Anadolu Genel dağılımı: Türkiye Bulunduğu kareler:A4 A5 A7 A8 A9 B8 B9 B10 C10

H. pastinacifoliumC. Koch in Linnaea 16:360 (1842). Syn: H. sisianensis Boiss. & Buhse, Aufz. Transk. Pers. 101 (1860). Map 71, p. 495.   Perennial, 15-90 cm, not aromatic, with a collar of broad flatfibres. Stems 3-6mm diam. below, slenderly sulcate, glabrous, pubescent or tomentose. Lower leaves 1-2-pinnate, lamina oblong to narrowly triangular, 5-30 x 3-13 cm, the primary * pinnae either ovate to suborbicular leaflets, 2-8 x l-4(-6) cm, obtuse to acuminate, dentate to incised, sometimes 2-3-lobed, or the lower pinnae 1 -pinnate with 3-5(-7) similar leaflets. Upper cauline leaves with sheaths scarcely dilated or very broadly so. Rays 5-23, subequal or very unequal, 2-12 cm. Bracteoles variable or none. Flowers ± white, the outer ± radiant. Ovary puberulent to glabrous. Fruits broadly oblong elliptic to obovate-orbicular, 5-9 x 3•5-5 mm, pubescent to glabrous, sometimes scabridulous; dorsal vittae clavate, ¼-⅔(- ¾ ) as long as mericarp; commissural vittae apparently absent. H. pastinacifolium is treated here as a very variable complex in which 3 weakly delimited, allopatric subspecies can be recognized in Turkey. 1. Sheaths of upper cauline leaves oblong-lanceolate, scarcely dilated; leaflets (or their segments) of lower leaves ovate-oblong, pubescent; rays very unequal, the longest 8-12 cm in fruit; dorsal vittae ½-⅔ as long as mericarp subsp. pastinacifolium 1. Sheaths of upper cauline leaves broadly dilated, truncate to retuse; rays usually subequal and shorter; leaflets of lower leaves suborbicular to ovate-lanceolate, glabrous, pubescent or tomentose; dorsal vittae usually ¼-½ (- ¾ ) as long as mericarp    2. Leaflets ovate-lanceolate, subacuminate, shortly pubescent subsp. transcaucasicum    2. Leaflets broadly ovate to suborbicular, glabrous, pubescent, shortly hirsute or tomentose subsp. incanum subsp. incanum (Boiss. & Huet) Davis, comb. et stat. nov. Syn: H. incanum Boiss. & Huet in Boiss., Diagn. ser. 2(2):93 (1854)! H. incanum Boiss. & Huet var. lazicum Boiss., Fl. Or. 2:1048 (1872)! H. cuneatilobatum Hausskn. & Bornm. in Feddes Rep. Beih. 89(2):287 (1941)! H. albowii Manden., Kavk. Heracleum 83 (1950); H. anatolicum Manden. in Notes R.B.G. Edinb. 24:179, f.l (1962)! Fl. 7-9. Rocky slopes, gullies, screes, summits, 1800-3500 m.   Type: [Turkey B8 Erzurum] in monte Tech Dagh supra Erzurum, 2338-2743 m, Huet du Pavilion (G! K!).   N. & E. Anatolia. A4 Zonguldak: Kel Tepe above Karabük, 1800 m, D. 38809! A5 Amasya: Ak Da., 1800 m, Bornm. 1890:2879 (type of H. cuneati-lobatum)! A7 Gümüşane: Karagoelldagh, Sint. 1894: 7270! A8 Bayburt: Kara Kaya Da., N. of Bayburt, 3505 m, Balls 1828! Rize: d. Ikizdere, Baltaş Tepe, 3200 m, D. 21110! Khabakor, 3300 m, Bal. 1384 (type of H. incanum var. lazi-cum)] Giresun: Balabandağlari (Kiline Tepe) above Tamdere, 2700 m, D. 20556 (type of H. anatolicum -dorsal vittae ¾ as long as mericarp)! B8 Bingöl: Bingöl, nr. Gumgum, 2980 m, Kotschy 1859: 386! B9 Bitlis: Süphan Da., 3300 m, D. 24748!   Endemic. Growing in drier, rockier habitats than the other two subspecies, and extremely variable in indumentum. Glabrous forms of subsp. incanum are confined to N. Anatolia (where they have been collected together with pubescent plants), and the hairiest forms are found in the more arid interior. The type of H. anatolicum is a form with exceptionally long vittae.