Apiaceae

Heracleum pastinacifolium subsp. pastinacifolium

Heracleum pastinacifolium subsp. pastinacifolium

Heracleum L. Heracleum pastinacifolium C. KOCH subsp. pastinacifolium C. KOCH Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 6 Son çiçeklenme zamanı: 7 Habitat: nemli çayırlık ve dere yakınları Minimum yükseklik: 2150 Maksimum yükseklik: 3500 Endemik: - Element: ? Türkiye dağılımı: GD. Anadolu Genel dağılımı: Ermenistan, Azerbaycan Bulunduğu kareler:A4 A5 A7 A8 A9 B8 B9 B10 C10

H. pastinacifoliumC. Koch in Linnaea 16:360 (1842). Syn: H. sisianensis Boiss. & Buhse, Aufz. Transk. Pers. 101 (1860). Map 71, p. 495.   Perennial, 15-90 cm, not aromatic, with a collar of broad flatfibres. Stems 3-6mm diam. below, slenderly sulcate, glabrous, pubescent or tomentose. Lower leaves 1-2-pinnate, lamina oblong to narrowly triangular, 5-30 x 3-13 cm, the primary * pinnae either ovate to suborbicular leaflets, 2-8 x l-4(-6) cm, obtuse to acuminate, dentate to incised, sometimes 2-3-lobed, or the lower pinnae 1 -pinnate with 3-5(-7) similar leaflets. Upper cauline leaves with sheaths scarcely dilated or very broadly so. Rays 5-23, subequal or very unequal, 2-12 cm. Bracteoles variable or none. Flowers ± white, the outer ± radiant. Ovary puberulent to glabrous. Fruits broadly oblong elliptic to obovate-orbicular, 5-9 x 3•5-5 mm, pubescent to glabrous, sometimes scabridulous; dorsal vittae clavate, ¼-⅔(- ¾ ) as long as mericarp; commissural vittae apparently absent. H. pastinacifolium is treated here as a very variable complex in which 3 weakly delimited, allopatric subspecies can be recognized in Turkey. 1. Sheaths of upper cauline leaves oblong-lanceolate, scarcely dilated; leaflets (or their segments) of lower leaves ovate-oblong, pubescent; rays very unequal, the longest 8-12 cm in fruit; dorsal vittae ½-⅔ as long as mericarp subsp. pastinacifolium 1. Sheaths of upper cauline leaves broadly dilated, truncate to retuse; rays usually subequal and shorter; leaflets of lower leaves suborbicular to ovate-lanceolate, glabrous, pubescent or tomentose; dorsal vittae usually ¼-½ (- ¾ ) as long as mericarp    2. Leaflets ovate-lanceolate, subacuminate, shortly pubescent subsp. transcaucasicum    2. Leaflets broadly ovate to suborbicular, glabrous, pubescent, shortly hirsute or tomentose subsp. incanum subsp. pastinacifolium. Fl. 6-7. Moist meadows and by streams, 2150-3500 m.   Described from Armenia.   S.E. Anatolia. B9 Van: 30 km from Başkale to Hoşap, 2600 m, D. 45920! B1O Van: 5 km N.E. of Başkale, 2150 m, D. 45862! C10 Hakkari: Mor Da., 2600-3500 m, Watson et al. 2965!   Armenia, Soviet Azerbaijan. Grossheim (Fl. Kavk. 7: map 132) records H. pastinacifolium from Ararat, but as H. transcaucasicum is included in Tamam-schian's synonymy, the identity of the record is doubtful. I am adopting Mande-nova's concept of H. pastinacifolium (sensu stricto) here, though C. Koch's original description says the leaflets are 'long-acuminate' - a feature of Turkish material assigned here to the following subspecies.