Asteraceae

Inula heterolepis

Inula L. Inula heterolepis BOISS. Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 6 Son çiçeklenme zamanı: 8 Habitat: kireçtaşı kayalık, uçurum ve çağıllık Minimum yükseklik: 0 Maksimum yükseklik: 1500 Endemik: - Element: D. Akdeniz Türkiye dağılımı: KB., B., ve G. Anadolu Sahil Genel dağılımı: G. Lübnan Bulunduğu kareler:A1 A2 A3 A4 A7 A9 B1 B2 B3 B4 B5 B6 B7 B9 B10 C1 C2 C3 C4 C5 C6

I. heterolepis Boiss., Diagn. ser. 2(3): 12 (1856). Syn: I. Candida (L.) Cass. var. heterolepis (Boiss.) Fiori in Ann. R. Istit. Sup. Forest. Naz. 9:41 (1924)! I. verbascifolia (Willd.) Hausskn. subsp. heterolepis (Boiss.) Tutin in Bot. J. Linn. Soc. 67:283 (1973)! Ic: Boul., Fl. Lib. Syr. t. 208 (1930). Map 5, p. 59.   Densely white-felted perennial herb with thick woody root-stock, covered with remains of petioles and often strongly lanate-pilose. Stems several, 15-40 cm, ± erect, branching above. Basal leaves ovate or elliptic-ovate, 2-6x1-3 cm,borne on petioles 1-7 cm long, obtuse or subacute at apex, attenuate at base, margins entire or obsoletely denticulate; cauline leaves similar, decreasing in size, upper ones ± sessile. Capitula disciform, 4-6 in racemes or up to 20 in subcorym-bose panicles. Involucre 0∙75-1 cm broad; phyllaries 4-5-seriate, imbricate; outer oblong, obtuse, 3-4x1-1-5 mm, white-felted; inner linear, acuminate, 5∙-5-7 x 0∙5 mm, pubescent. Female flowers narrowly tubular and subligulate, (2-)3-3∙75(-5) mm. Disc flowers 5-6 mm. Achenes 1∙5-2 mm, finely pubescent in upper half. Pappus white, 4∙5-5∙5 mm. hairs 10-15, scabrous, free at base. Fl. 6-8. On limestone rocks, cliffs and screes, s.1.-1500 m.   Syntypes: [Turkey Bl & Cl] in Sipylo supra Magnesiam, Boissier (BM!); in Caria, Pinard; ad Smyrnam (Izmir). Balansa 246 (K!).   Mainly N.W., W. & S. Anatolia, usually near the coast. A2(A) Bursa: Olympus (Ulu Da.), vi 1855, Kirk] Kocaeli: Gebze to Izmit, 50 m, D. 36249! Bl Izmir: Smyrna (Izmir), vi 1827, Fleischer] B2 Kütahya: Gediz to Kütahya, 1000 m, D. 36914! Cl Aydin: Samsun Da. at Priene, 100-900 m, D. 18343! C2 Antalya: Elmali. Bourgeau 154! Muğla: between Muğla and Ulu, D. 13486! Denizli: between Çukur Köy and Kizilhisar, D. 13280! C3 Antalya: d. Kemer, Teke Da. nr Ovacik, 1200 m, D. 15218! Isparta: Anamas Da., 1500 m, Sorger 68-37-28a! C4 Konya: Ermenek, 1300-1400 m, D. 16154! Antalya: Alanya, D. 14480! Içel 6 km W. of Mut, Sorger 66-22-2! C5 Içel: 16 km E. of Silifke, Sorger 71-22-3! Adana: Pozantı, Toros Da., 1100 m, Aberdeen Univ. Amanus Exped. Al 25! Is: Lesvos, Rech, 1273; Khios, Platt 345! Ikaria, Barbey 920! Samos, Bothmer & strid21813; Fournoi, Bothmer & Strid 21837; Lipsoi, Runemark; Leros, d'Urv.; Kalimnos, Fors.-Maj. 558; Kos, Rech. 8046; Tilos, Ade; Simi, Gathome-Hardy 52! Khalki, Gathorne-Hardy 779! Rodhos, Bourgeau 93!   Some records in literature, especially those from N. Anatolia, refer to I. aschersoniana or to I. fragilis.   S. Lebanon. E. Medit. element. This is the commonest of the Turkish species that belong to the I. Candida complex (/. heterolepis, I. aschersoniana, I. fragilis and I. anatolicd). I. Candida L. is itself endemic to Crete but the species, or sub-species as they are sometimes treated, closely related to it are widespread throughout the Eastern Mediterranean.   Leaves are generally almost equally tomentose on both surfaces but sometimes, as in Sorger 71-22-3 (C5 Içel), they may be weakly tomentose and green on the upper surface. Female corollas are also variable in length; usually they are about 3∙5 mm but in Eig & M.Zohary (C5 Içel: nr Goezne, 1000-1100 m, 16 viii 31) they are 2 mm, whilst, in the above Sorger specimen, they measure 5 mm. The corollas are also variable in their lobing, sometimes ending obliquely, sometimes with well developed teeth. Generally there are about 20 female flowers in each capitulum but particularly in the south (Muğla and Antalya) there is a trend in which the number is commonly reduced by half, then further to 1-2, e.g. in D. 14093 (C3 Antalya: d. Kemer, Teke Da. between Kuzdere and Gidelma, 800 m) leading finally to the complete absence of female flowers in D. 15041 and 15149 (both from C3 Antalya: d. Kemer, Kesme boğaz and Gönük, 0-100 m). These latter specimens are quite distinct from I. fragilis not only geographically but are taller plants with narrower capitula.