Lamiaceae

Lamium garganicum subsp. reniforme

Lamium L. Lamium garganicum L. subsp. reniforme (MONTBRET ET AUCHER EX BENTHAM) R. MILL Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 4 Son çiçeklenme zamanı: 9 Habitat: gölgeli kalkerli ve volkanik kayalar kaya yarıkları, çağıllık, duvarlar, harabeler, nehir Minimum yükseklik: 400 Maksimum yükseklik: 2300 Endemik: - Element: ? Türkiye dağılımı: Karasal ve G. Anadolu Genel dağılımı: Kafkasya, KB. İran, K. Irak Bulunduğu kareler :A1 A2 A3 A4 A5 A7 A8 B2 B3 B4 B5 B6 B7 B8 B9 C3 C4 C5 C6 C7 C8 C9 C10

L. garganicum L., Sp. PI. ed. 2: 808 (1763). Perennial, often sprawling over rocks, sometimes forming mats. Stems 6- 45 cm, villous to glabrous. Leaves broadly ovate to reniform, lamina 3-52 x 3- 46 mm, crenate or serrate, villous to ± glabrous. Verticillasters 1-5, (2-)4-8(-12)-flowered. Bracteoles 3-9(-10.5) mm. Calyx (6-)8-18(-20.5) mm, tube glabrous to pubescent or sparingly villous, 5-10(-l 1) mm; teeth 2-9 mm. Corolla purplish-pink (rarely white), with purple-striate tube and usually a large purple blotch on lower lip, 22-40(-45) mm; tube straight (rarely curved), without annulus, 14-29 mm; upper lip 6-16 mm, deeply or shortly bifid, emarginate, retuse or ± entire, the lobes themselves sometimes with notched to bifid apices; lower lip 6-20 mm, lateral lobes without or occasionally with a short subulate appendage. Nutlets 2.5-3.5 x 1.3-1.6 mm, blackish-green or brown, usually with a few paler spots. 1. Corolla white with purple veins and bluish-purple blotches at tips of both lips; tube slightly curved; plant mat-forming, with very densely whitish-villous leaves g. subsp. pulchrum 1. Not as above    2. Stems villous      3. Glandular only in inflorescence b. subsp. reniforme      3. Glandular throughout         4. Corolla 34-40 mm f. subsp. lasioclades         4. Corolla (20-) 2 5-3 2 mm             5. Calyx teeth subequal to tube b. subsp. reniforme             5. Calyx teeth shorter than tube e. subsp. rectum    2. Stems glabrous to densely pubescent                6. Upper lip of corolla deeply bifid, lobes 3-5.5 mm                   7. Bracteoles 4-7 mm a. subsp. striatum                   7. Bracteoles c. 3 mm b. subsp. reniforme                6. Upper lip of corolla shortly bifid to subentire, lobes 0-2.5 mm                       8. Hairs of stem more than 0.5 mm b. subsp. reniforme                       8. Hairs of stem 0-0.5 mm                         9. Median leaves longer than 40 mm                           10. Leaf incisions 5-5.5 mm deep a. subsp. striatum                           10. Leaf incisions 1-4.5(-5) mm deep                                11. Erect (N.W. Turkey) d. subsp. laevigatum                                11. Sprawling or ascending (C. & S. Anatolia) b. subsp. reniforme                         9. Median leaves less than 40 mm                                  12. Leaves dentate or acutely crenate c. subsp. nepetifolium                                  12. Leaves obtusely crenate                                    13. Calyx teeth (2/3-)3/4-9/l Ox tube b. subsp. reniforme                                    13. Calyx teeth (1/3-) 1/2-2/3 X tube                                      14. Stems pubescent, hairs 0.2-0.3 mm a. subsp. striatum                                      14. Stems ± glabrous or puberulent, hairs (if any) usually less than 0.1 mm, never exceeding 0.2 mm                                         15. Sprawling or decumbent; leaves sometimes broader than long (C, S. & E. Anatolia) b. subsp. reniforme                                         15. ± Erect; leaves ± longer than broad (N.W. Turkey only) d. subsp. laevigatum a. subsp. striatum (Sm.) Hayek, Prodr. F1. Balc. 2:275(1929). Syn: L. striatum Sm. in Sibth. & Sm., Prodr. Fl. Graec. 1:405 (1809); L. grandiflorum sensu Sieber, Avis 5 (1821) non Pourr. (1788); Dracocephalum lamiifolium Desf., Choix P1. 33: t. 15 (1808)! Ic: Sibth. & Sm., Fl. Graeca 6: t. 557 (1827). Map 22. Fl. 3-5. Shady banks, volcanic cliffs, schist, 390-900 m. Type: [Greece & Aegean] in ruderatis Graeciae et Archipelagi copiose, Sibthorp (holo. OXF). Islands. Is: Lesvos, Fili a village, 380-450 m, Stamatiadou 4727! ibid., mt. Olympos, 500 m, Edmondson & McClintock 2169! ibid., 700-900 m, Stamatiadou 4694! ibid., Vatoussa, Rech. 5842! Samos, S. of Vurliotes, Runem. &Snog. 19907! Greece, Aegean. E. Medit. element. b. subsp. reniforme (Montbret & Aucher ex Bentham) R. Mill in Notes R.B.G. Edinb. 38:34 (1980). Syn: L. reniforme Montbret & Aucher ex Bentham in Ann. Sci. Nat. (Paris) ser. 2, 6:47 (1836)! L. striatum Sm. var. reniforme (Montbret & Aucher ex Bentham) Boiss., Fl. Or. 4:757 (1879)! L. striatum Sm. var. glabrum Beg. & Diratz., Contr. Fl. Armen. 93, t. 12 f. 6-8 (1912). Map 22. Fl. 4-9. Shady limestone and igneous rocks, crevices, scree, walls, ruins, riverbanks, 400-2300 m.   Type: [Turkey A8 Trabzon/Gümüşane] in montibus regionis Lagisthan inter Baihout (Bayburt) et Trebizondem (Trabzon), Montbret & Aucher (holo. P, iso. G! K!).   Inner & S. Anatolia. A4 Ankara: Ayaş Da., 1350 m, Akman 3086! A5 Amasya: Ak Da., E. of Kocaciktepe, 1500 m, Alpinar (ISTE 36943)! A7 Gümüşane: Bayburt Osluk Köprüsü, 1750 m, Tobey 2139! A8 Çoruh: Kordevan Da. (Yalnizçam Daglari) nr Kütül Y., 200° m, D. 30184! B2 Kütahya: nr Kütahya, 800 m, Balls 633! B5 Kayseri: Bakir Da. at Akoluk Y. above Kisge, 1900 m, D. 19447! B6 Maraş: Binboga Da. above Yalak, 2300-2400 m, D. 20123! B7 Erzincan: Eğin (Kemaliye), Szanduk (Sandik), Sint. 1890:2390! B8 Bitlis: Kambos Da., 1830 m, D. 23491! B9 Bitlis: Bitlis, 1400 m, Frödin 1939:261 (as L. striatum var. nepetaefolium)! C3 Antalya: Termessos, Güllük Da., 850-1000 m, Bozakman & Fitz 1970:26! C4 Içel: d. Anamur, Çamurla Y. to Olucak, 2000 m, D. 16295! C5 Adana: d. Karaisali, Koca Çukur Y., N.W. of Pozanti, 1750 m, Hub.-Mor. 15607! C6 Hatay: Nur Da. 16 km S.W. of Yarpuz, 1670-1850 m, Buttler & Erben 17835! C8 Mardin: Mardin castle, 1200 m, D. 28339! C9 Hakkari: nr Hakkari, 1800 m, 21 v 1966, Eiselt! C10 Hakkari: nr Ziri, S. of Cilo Da., 1830 m, Trelawny 1840!   Caucasia, N.W. Iran, N. Iraq. The most widespread subspecies of L. gar-ganicum in Turkey, showing considerable variation especially in leaf morphol-. ogy. Local populations, e.g. in Kayseri, Mardin, Van and Bitlis vilayets, are often recognisably distinct but, because variation is ± continuous throughout the range of the subspecies, these variants are not formally recognised here. Forms from S.W. Anatolia (B3 Isparta: Sultan Da., 1600 m, Sorger 67-2-5! C3 Isparta: Çiçek Da. 1300-1600 m, Sorger 67-5-54! 16 km N.E. of Eğridir, 980 m, Buttler & Uzunoğlu 12787!) have some characters of subsp. reniforme but differ by their more triangular, sometimes more acutely toothed leaves; they appear to be transitional to L. lycium. Some specimens from N. & C. Anatolia (A3 Bolu: Mudurnu, 1500 m, Akman 6121! A5 Çorum: Haciköy to Osmancik, 800 m, Tobey 2564! B3 Konya: Saray Da., 1600 m,v 1902,Zeder-bauer! B4 Afyon: Bayat, Koroğlu Tepe, 1350 m, Vural 310!) have small, triangular, ± dentate laeaves and resemble subsp. nepetifolium from S. Anatolia. Two specimens from C5 Nigde (Hasan Da. nr Taşpinar Y., 2000 m, D. 18995! ibid., 2600 m, D. 18962!) resemble subsp. reniforme in leaf morphology but differ from all other gatherings by their densely pubescent-pilose (almost villous) stems and leaves. They differ from each other in corolla morphology: D. 18995 has the upper lip and lateral lobes entire, while in D. 18962 the upper lip is bifid to 2.5 mm and the lateral lobes have a 0.5 mm' appendage. They may represent one or two new taxa but further collections from the area are needed.