Scrophulariaceae

Linaria genistifolia subsp. genistifolia

Linaria genistifolia subsp. genistifolia

Linaria MILLER Linaria genistifolia (L.) MILLER subsp. genistifolia (L.) MILLER Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 5 Son çiçeklenme zamanı: 8 Habitat: koruluk, çalılık, maki, kayalar, topraklı yamaçlar, yol kenarları Minimum yükseklik: 0 Maksimum yükseklik: 2100 Endemik: - Element: Avrupa-Sibirya Türkiye dağılımı: K. Türkiye, Seyrek Olarak Başka Yerlerde Genel dağılımı: O. ve GD. Avrupa Doğusunda Sibirya, Kafkasya Bulunduğu kareler :A1 A2 A3 A4 A5 A6 A7 A8 A9 B1 B2 B3 B4 B5 B6 B7 B8 C2 C3 C4 C5 C6 C8

L. genistifolia (L.) Miller, Gard. Dict. ed. 8, no. 14 (1768); Alex in Can. J. Bot. 40:295-307 (1962).   Perennial herb, very variable. Stems 20-130 cm, ascending or erect, simple or paniculate. Leaves alternate. ± dense, median ovate, lanceolate or linear, 15-60 x 0.5-20 mm, rarely subcordate at base, 1-3-nerved. Racemes dense to lax and elongate. Bracts lanceolate, seldom reaching top of calyx, lowest sometimes deflexed. Lower pedicels 2-4(-6) mm. Calyx lobes ovate to narrowly linear-lanceolate, rounded at base, acute, herbaceous throughout, 2-5. mm, always glabrous. Corolla bright or sulphur yellow, with or without reddish venation, (6-)8-12(-14) mm; spur conical-subulate, (3-)5-8(-10) mm, ± straight. Capsule spherical, 3-4 mm. slightly longer to shorter than calyx. Seeds compressed-tetrahedral, 1-1.2 mm, rugulose, angles often narrowly winged. Fl. 5-8. A narrower circumscription of the polymorphic L. genistifolia is adopted here than in Fl. Europaea 3:230 (1972), treating both L. grandiflora Desf. and L. sofiana Velen. as separate species. Within L. genistifolia, six weakly delimited subspecies can be recognized in Turkey. 1. Median leaves 10-45 x longer than broad, linear-lanceolate to narrowly linear. 1-4 mm broad, 1-nerved; stems ascending to erect, 20-60 cm, usually arising from a creeping stock    2. Flowers bright brassy yellow (lutea); median leaves linear to linear-lanceo-, late, flat, (l-)2-4 mm broad subsp. linifolia    2: Flowers palish yellow (flava); median leaves linear-filiform, usually involute, 0.5-1 (-1 .5) mm broad subsp. polyelada 1. Median leaves 2-10 x longer than broad, ovate to lanceolate, 3-20 mm broad. l-3-nerved; stems always erect (unless grazed), 23-130 cm      3. Flowers clear yellow, not reddish-veined        4. Corolla bright yellow (lutea); inflorescence ± lax. at least in fruit; calyx lobes ovate-lanceolate to lanceolate subsp. genistifolia        4. Corolla paler yellow (flava); inflorescence dense, even in fruit (though often long); calyx lobes usually lanceolate to linear-lanceolate subsp. confertiflora      3. Flowers yellow with reddish-brown venation          5. Calyx lobes lanceolate, 3-5 mm; corolla 10-12 mm, tube at least as broad as long; spur only 5-7 mm subsp. praealta          5. Calyx lobes ovate, 2-2.5 mm; corolla 8-9 mm, tube narrower than long; spur c. 8 mm subsp. arfvinensis subsp. genistifolia. Syn: Antirrhinum genistifolium L., Sp. PI. 616 (1753)! Linariu euxina Velen. in Bot. Centr. 36: 125 (1888); L. politico Kupr. in Trudy Bot. InSt. Akad. Nauk SSSR 9:64 (1950). Ic: Sibth. & Sm., FI. Graeca 6: t. 596 (1827); Fl. RPR 7: t. 78 f. 1 (I960). Woodland, scrub, macchie, rocks, earthy slopes, roadsides, s./.-21OO m.   Described from Siberia. Lectotype: Hb. Linn. 767/40!   Most common in N. Turkey, but scattered elsewhere. A1(E) Kırklareli: Istranca Da., Demirköy to Iğneada, Kayacık 32! A2(E) Istanbul: Sariyer, 200 m, Edmondsok 434!. A2( A) Istanbul: Büyük De., 23 v 1897, Nemetz! A3 Zonguldak: coast S.W. of Alapli, D. 37457! A5 Amasya: Merzifon, 1904, Maniss.! A6 Ordu: Çambasi Yavuz Bükü, 2100 m, Tobey 1422 (inflorescence dense)! A7 Trabzon: Sumila, Sint. 1889:1729 (large shade form)! B2 Kutähya: c. 35 km W. of Abide on road to Simav, 900 m, Coode & Jones 2618! B6 Sivas: Çamlibel Da., 1600-1700 m, 14 vii 1939. Reese! B7 Elaziğ: Kekan at Murad su, Sint. 1889:402! C4 Içel: 21 km from Gilindire to Gülnar, 670 m, Hub.-Mor. 10379! C. & S.E. Europe eastwards to Siberia; Caucasia? Euro-Sib, element. L. euxina Velen. may be a dwarf (often grazed) small-leaved maritime ecotype. The Crimean L. politico Kupr. is only a small-flowered variant of subsp. genistifolia and in Anatolia falls within the morphological range of the latter. L. iberica Kem.-Nath. (Fl. Gruzii 7:504, 1952) and L, imeretńica Kem.-Nath. (op. cit. 503) are also very close to subsp. genistifolia and certainly conspecific. subsp. confertiflora (Boiss.) Davis in Notes R.B.G. Edinb. 36:6 (1978). Syn: L. monochroma Boiss. & Heldr. in Boiss., Diagn. ser. 1(12):41 (1853)! L. genisti-folia (L.) Miller var. confertiflora Boiss., Fl. Or. 4:377 (1879)! Figure 21, p. 655. Dry rocky slopes (calcareous and igneous), phrygana, pastures, fallow fields, 200-2000 m. Syntypes: [S.W. & C. Anatolia] in campis Pisidiae et Isauriae rara, circa [C3 Konya] Beycheher (Beyşehir), Heldreich (G!); [C3 Isparta] Eğirdir (Eğridir). Heldrcich; [C4 Konya] Koniah, vi. Heldreich (syntypes of L. monochroma. iso. BM! E!); [C2 Antalya] Elmalu, Botirgeau [186] (iso. G!). Mainly Inner & S. Anatolia; common. A4 Ankara: Kizilcahamam to Cam-koru, Alpay 261 ! A8 Gümüşane: Gümüşane, 1 100 m. Balls 174! B2 Izmir: Boz Da., 1300 m, Sorger 68-13-10! B3 Konya: Akşehir, Bornm. 1899:5395! B4 Ankara: 40 km E. of Ankara, 18 iv 1962, Andres! B6 Maraş: Binboğa Da., 1900 m, D. 20037! B7 Tunceli: Munzur Da., 2000 m, D. 31314! B8 Erzurum: mts. between Ilica and Tercan, D. 30908! C2 Denizli: Honaz Da., 1200 m, A.Baytop & E.Tuzlaci. C3 Burdur: 20 km S. of Burdur, Alava 5206! C4 Konya: E. of Ermenek, 1250 m, Sorger 66-32-23! C5 Konya': Cilician Gates (Gülek Boğazi), Bal. 1855:684! Adana: Nur Da. (Amanus), 16 km S.W. of Yarpuz, 1850-2000 m, Buttler & Erben 17842! Endemic? Ir.-Tur. element. Intergrading with subsp. genistifolia and subsp. linifolia. subsp. linifolia (Boiss.) Davis in Notes R.B.G. Edinb. 36:7 (1978). Syn: L. concolor Griseb., Spic. 2:21 (1844); L. syspirensis C. Koch in Linnaea 22:717 (1849); L. genistifolia (L.) Miller var. linifolia Boiss., Fl. Or. 4:378 (1879)! Ic: Stoj. & Stef., Fl. Bulg. 996 (1925), as L. concolor. Mainly rocky mountain slopes and screes, 1400-2700 m. Lectotype (Davis, op. cit. 7, 1978): [Turkey A8 Rize] valle Djimil (Cimil), 1830 m, [1866], Balansa (holo. G!). Scattered in N.W. & Inner Anatolia. A2(A) Bursa: hills nr Bursa, Pichler 225! A4 Kastamonu/Çankiri: İlgaz Da., 1400 m. Khan et al. 675! A5 Kastamonu: Tosya, Dalscha Dagh (nr Çakirlar), Sint. 1892:4696! A7 Giresun: Balaban Da. (Kilinç Tepe), 2700 m, D. 20601! Gümüşane: Stavros, 2130 m. Balls 517! A8 Erzurum: 30-32 km from Oltu to Tortum, 2100-2200 m, D. 47622! A9 Kars: 11 km W. of Sarikamiş, 2260 m, Buttler 14504! B1 Balikesir: mt. Ida (Kaz Da.), Sint. 1883:732! B2 Izmir: Bozdağ Y., 18 vii 1854, Bal.! B5 Kayseri: Argaeus (Erciyas Da.), Bal. 1856:527! B7 Tunceli: Pülümür to Mutu, nr pass, 1450 m, 26 vi 1951, Reese! C2 Denizli: Honaz Da., Karbadiç, 2000 m, A.Baytop & E.Tuzlaci. C5 Adana: Karsanti, Karataş Çam oluğu, Yurdakulol 62 (doubtful)! Balkans, Crimea, Caucasia. Variable as delimited here, and perhaps poly-phyletic within the species. A peloric form has been seen from A7 Gümüşane (Jenkins 2303!). subsp. praealta (Boiss.) Davis in Notes R.B.G. Edinb. 36:7 (1978). Syn: L. praealta Boiss. & Bal. in Boiss., Diagn. ser. 2(3): 162 (1856)! L. balansae Boiss. & Reuter in Boiss., Diagn. ser. 2(3): 163 (1856), grazed! L. genistifolia (L.) Miller var. venosa Boiss., Fl. Or. 4:378 (1879) p.p.! Quercus scrub, limestone slopes, stream beds, chalky vineyards, 800-2000 m. Type: [Turkey C5 Içel] in fauce Güzel Dere [Güzel De.] supra pagum Kech-lick (Keklik) non procul a Mersina in Cilicia littorali, vi [1855], Balansa (holo. G!). S. & S.E. Anatolia. C5 Adana: Bürücek, 1130 m. Balls 1118a! C5/6 Hatay: Amanus, Haradj. 411! C6 Gaziantep: Nurdaği pass, W. of Fevzipaşi, 1150 m. Sorger 71-37-18! Maraş: Ahir Da.. 900 m. Balls 1118! C8 Mardin: Mardin vineyards, D. 28372! Endemic. E. Medit. element. The corolla is proportionally broader than usually found elsewhere in L. genistifolia, but as this feature occurs sporadically in other parts of the species' range, specific status is not justified. subsp. polyclada (Fenzl) Davis in Notes R.B.G. Edinb. 36:7 (1978). Syn: L. polyclada Fenzl in Tchihat., Asie Min. Bot. 2:23 (1860)! L. genistifolia (L.) Miller var. venosa Boiss., Fl. Or. 4:378 (1879) p.p.! Volcanic tuff, fallow fields, 800-1240 m. Type: [Turkey C5 Içel] Ciliciae Bulgardagh, inter segetes prope pagum Gulek, 1235 m, Kotschy 1836:399 (iso. E! G!). S. & C. Anatolia. B5 Nevşehir: Ürgüp to Nevşehir, Alava 7094! Göreme valley, 1100 m, 21 viii 1972, Shmida & Luria! C5 Niğde: mt. above Maden, 1700 m, Siehe 1908:299! Endemic. Ir.-Tur. element. The type of L. polyclada Fenzl has a shorter spur (3 mm) than material from Nevşehir (4-6.5 mm). subsp. artvinensis Davis in Notes R.B.G. Edinb. 36:7 (1978). Syn: ?L. adzharica Kem.-Nath. in Zam. Sist. Geog. Rast. 23:40 (1963). Rock crevices, dry roadsides. 400-600 m. Type: Turkey A8 Çoruh: Artvin, 400 m, Stainton 8290 (holo. E!). N.E. Anatolia; rare. A8 Çoruh: 600 m, Akeroyd 188! Endemic? Euxine element. Perhaps closest to subsp. genistifolia, despite its reddish-veined flowers.