Sanguisorba minor subsp. minor

Rosaceae

Sanguisorba minor
subsp. minor

Sanguisorba L.
Sanguisorba minor SCOP. subsp. minor SCOP.
Ömür: Çok yıllık
Yapı: ot
Hayat formu:
İlk çiçeklenme zamanı: 0
Son çiçeklenme zamanı: 0
Habitat: ?
Minimum yükseklik: 10
Maksimum yükseklik: 2350
Endemik: -
Element: ?
Türkiye dağılımı: B. Türkiye, KB. ve Karasal Anadolu
Genel dağılımı: Avrupa, K. Afrika, GB. ve O. Asya
Bulunduğu kareler :A1 A2 A5 A7 A8 A9 B1 B2 B3 B4 B5 B6 B7 B8 B9 C1 C2 C3 C4 C5 C6 C7 C8 C9 C10

S. minor Scop., Fl. Carn. 110 (1772). Figure 1, p. 87.
  Stems erect, (5-) 10-75 cm, glabrous, puberulous or sparsely pilose, sparingly branched. Lower leaves in rosettes at the base of the stems, ± oblong, pinnate with (7-) 11-17 orbicular to oblong to ovate, deeply serrate leaflets, usually pilose or sericeous-pilose beneath. Upper leaves few, similar to rosette leaves but with fewer, relatively narrower leaflets. Heads ± globose. Sepals green with whitish or pinkish margins, ovate-oblong, 2-3 mm. Hypanthium forming a hard, nut-like 4-angled fruit with hairy, wrinkled or verrucose faces.
  A very variable species which Nordborg divides into 6 subspecies, definable on fruit characters, which are not correlated with any other vegetative or floral features. Of the 6 subspecies, 4 occur in Turkey.
1. Fruit puberulous, obovoid or ovoid, rounded to the apex, not compressed, the 4 wings obscure, the surface wrinkled or almost smooth, 2-3 x 1-1.5 mm (Figure 1) . subsp. lasiocarpa
1. Fruit glabrous, rounded or ± acute at the apex, compressed or not, wings often conspicuous, surface usually conspicuously wrinkled or verrucose, 2.5-4 x 1.3 mm
   2. Angles not prominently winged, rather flattened; fruit fusiform, somewhat compressed, surface wrinkled, 3-3.7 x 1-1.5 mm (Figure 1) subsp. minor
   2. Angles prominently winged, or if not, fruit markedly verrucose with conspicuous projections, not flattened, ovoid to obovoid
     3. Surface of fruit thrown into coarse folds and wrinkles; fruit 3-4 x1.6-2.5 mm (Figure 1) subsp. muricata
     3. Surface of fruit drawn up into prominent projections; fruit 3.5-4.5 x 2-3 mm (Figure 1) subsp. magnolii
subsp. minor. Syn: S. minor Scop., Fl. Carn. 110 (1772) sensu stricto; Poterium sanguisorba L., Sp. P1. 994 (1753). Ic: Bonnier, Fl. Comp. Fr., Suisse & Belg. 4: t. 183 (1920); Ross-Craig, Draw. Brit. P1. 9: t. 11 (1956). Type: Spain, Loefling 613 (Hb. Linn. 1124/1, photo!). W. Turkey, S.W. & Inner Anatolia, Islands. A1(E) Çanakkale: 25 km N. ofGelibolu, 10 m, Sorger 68-2-82! A8 Erzurum: nr. Tortum, Huet. B4 Ankara: between Kayaş and Kubaras, D. 13124! B5 Kayseri: Develi, D. 19168! B6 Sivas: 28 km N.W. of Divriği, 1500 m, Sorger 69-42-130! B7 Malatya: Gürün to Darende, 1700 m, D. 21864! Erzincan: 66 km from Suşehri to Erzincan, 1850 m, Apold et al. 200! B8 Erzurum: Palandöken Da., 2350 m, D. 47383! B9 Bitlis: Tatvan, 1900 m, D. 22356! C1 Izmir/Aydin: Samsun Da. {Nordborg 1967: f. 20). C2 Antalya: Ak Da., 1400 m, Sorger 65-26-12! C8 Mardin: Kasmi, Sint. 1888: 1104! Is: Samos, Nordborg 8249.
  Europe, N. Africa, S.W. & C. Asia. 2n = 28 (European material).
subsp. magnolii (Spach) Briq., Prodr. F1. Corse 2(1): 209 (1913). Syn: Poterium polygamum Desf., Cat. P1. Horti Paris. ed. 3:292 (1829); P. verrucosum Ehrenb. ex Decne. in Ann. Sci. Nat. ser. 2, 3:263 (1835); P. magnolii Spach in Ann. Sci. Nat. ser. 3, 5:38 (1846). Ic: Coste, Fl. Fr. 2:59 (1903). Fr. 4-6. Waste ground, fields, slopes, s.l.-1100 m.
Lectotype: [Algeria] Oran, Dıırieu (P, fide Nordborg 1967). W., S. & E. Anatolia, Islands. A2(A) Çanakkale: Renkoei (Erenköy), Sint. 1883:59! B1 Izmir: above Burnabat, Boiss. B2 Uşak: 8 km from Uşak to Güre, 800 m, Coode & Jones 2540! C1 Aydin: Kuşadasi, Nordborg 8268. C2 Denizli: Kazik Beli, 1225 m, Nordborg 8541. C3 Antalya: W. of Kale, Hub.-Mor. 10557. C4 Içel: 25 km from Mut to Silifke, Coode & Jones 1043! C5 Içel: Tarsus, 150 m, D. 26442! C6 Hatay: 8 km from Belen to Antakya, 600 m, Coode & Jones 2540! C7 Urfa: 43 km from Urfa to Viranşehir, 650 m, D. 42336! C8 Mardin: 11 km from Midyat to Gerçüş, 1100 m, D. 42928! Is: Lesvos, Rech. 1274; Khios, Rech. 5268! Samos, Rech. 3713! Ikaria, Rech. 4284; Rodhos, Mt. Profeta, 600 m, Rech. 1\A\!
S. Europe, N. Africa, S.W. Asia. Nordborg reports 2n — 28 for Turkish material.
subsp. muricata (Spach) Briq., Prodr. Fl. Corse 2(l):210 (1913). Syn: Poterium muricatum Spach in Ann. Sci. Nat. ser. 3, 5:36 (1846); P. gaillardotii Boiss., Diagn. ser. 2(2): 52 (1856); Sanguisorba gaillardotii (Boiss.) Hayek, Prodr. Fl. Balc. 1:699 (1926). Ic: Coste, Fl. Fr. 2:59 (1903). Fr. 6-7. Fields, forests, slopes, '250-2100 m.
Lectotype: [France] Aulnay, Spach (P, fide Nordborg 1967). Widespread. A1(E) Kirklareli: Pinarhisar to Erenlerköy, 250 m, A. Baytop 15823! A2(A) Bursa: Mudanya, Nordborg 8515. A2(E) Istanbul: Gümüş-pinar to Karacaköy, A. Baytop & Attila 15823! A4 Ankara: 26 km N. of Kizil-cahamam, 1300 m, Sorger 70-5-82! A5 Amasya: Amasya, 1100 m, Tobey 1189! A6 Ordu: Fatsa to Aybasti, 400 m, Tobey 1335! A7 Giresun: 25 km S. of Giresun, 500 m, Sorger 69-23-46! A9 Kars: c. 50 km S. of Kars, 1540 m, M. Zohary & Plitm. 2267-22! B1 Balikesir: Mt. Ida (Kaz Da.), nr. Juruk Obassi, Sint. 1883:939b! B2 Kütahya: between Kütahya and Gediz, 1100 m, M. & D. Zohary 6022! B3 Eskişehir: nr. Eskişehir, Birand & M. Zohary 3462! B4 Ankara: Dikmen, 19 vi 1953, M. Zohary! B5 Kayseri: Erciyas Da., 1400 m, Zederbaııer. B7 Erzincan: 34 km W. of Erzincan, 2060 m, 8 vii 1963, M. Zohary! B8 Erzurum: Chnys-kale (Hinis Kale), 1680 m, 18 vii 1916, Sapozhnikov. B9 Van: Van, 1720 m, D. 44656! C1 Aydin: Samsun Da., Nordborg 8271. C2 Muğla: Sandras Da., Köklüçam, D. 13607! C3 Isparta: Sütçüler, 1000 m, D. 15889b! C4 Konya: 20km W. of Konya, 1250 m, Nordborg 9841. C5 Içel: 1 km S.W. of Tekir Tepesi, 1300 m, Nordborg 8537. C6 Hatay: Belen, 800 m, Akman. C9 Hakkari: 5 km from Hakkari, Kaval, 1850 m, D. 45403! Is: Lesvos, nr. Ajassos, 400-700 m, Rech. 5610a; Samos, Leka, Nordborg 8254; Rodhos, Hedenborg.
Europe, N. Africa, S.W. Asia. Nordborg reports 2n = 28 from a number of collections from western Anatolia, and 2n — 56 for her collection from Bursa.
subsp. lasiocarpa (Boiss. & Hausskn.) Nordb. in Op. Bot. (Lund) 11:66 (1966). Syn: Poterium villosum Sm. in Sibth. & Sm., Prodr. F1. Graec. 2:238 (1816); P. lasiocarpum Boiss. & Hausskn. in Boiss., F1. Or. 2:733 (1872); Sanguisorba lasiocarpa (Boiss. & Hausskn.) Hand.-Mazz. in Ann. Nat. Hofmus. Wien 27:67 (1913). Ft: 7-9. Damp meadows, stream sides, 300-2400 m.
Lectotype: [Iran] ad Bebehan Persiae australis, Haussknecht (G, fide Nordborg 1967).
N.W., C. & E. Anatolia. A2(A) Balikesir: Marmaris Adasi, A. Baytop 13644! B3 Eskişehir: Kopet Da., Nordborg 8110. B7 Sivas: Divriği, 1200 m, Sorger 70-31-27! B8 Bitlis: Meleto Da., Sassun, 1500-2400 m, Hand.-Mazz. 2782! B9 Bitlis: Pelli Da., 1900 m, D. 46041! C7 Urfa: 11 km W. of Ceylanpinar, 300 m, Hub.-Mor. 15122. C8 Mardin: Mardin, Sint. 1888:1104! C9 Hakkari: Zap gorge, 55 km S. of Başkale, D. 23844! C10 Hakkari: 10 km from Yüksekova, Gevar Ovasi, 1950 m, Duncan & Tail 148!
Italy, Greece, S.W. Asia. Nordborg reports 2» = 56 for 2 Turkish collections from Istanbul (Aydos Da.) and Eskişehir (Kopet Da.).
Intermediates occur between all the subspecies. These are fully discussed by Nordborg (1967), who also gives a complete set of illustrations and distribution maps.
The following records are from Provinces or Squares for which. there are no records of the individual subspecies but from which flowering material (that cannot be identified to subspecies) is known: A1(E) Tekirdağ: Tekirdağ to Malakara, 3-50 m, D. 34690! A2(A) Bilecik: 2 km S. of Bilecik station, 400 m, D. 42076! A5 Yozgat: Akdağmadeni to Büyük Nalbant Da., 1800 m, Coode & Jones 2093! Samsun: Samsun, 5 m, Tobey 260! A7 Gümüşane: Tegrulun Köprü, Basi Köy, 950 m, Tobey 201! A8 Çoruh: Borçka, 200 m, Stainton 8205! B2 Manisa: Salihli, 800 m, Coode & Jones 2809! B7 Diyarbakir: 5 km N. of Ergani, 1000 m, D. 29012! B7 Tunceli: Tunceli to Pülümür, 1100 m, D. 29231!