Dipsacaceae

Scabiosa columbaria subsp. columbaria var. columbaria

Scabiosa L. Scabiosa columbaria L. subsp. columbaria L. var. columbaria L. Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 6 Son çiçeklenme zamanı: 9 Habitat: yolkenarları, çorak yerler, kayalık yamaçlar Minimum yükseklik: 0 Maksimum yükseklik: 2600 Endemik: - Element: ? Türkiye dağılımı: K. Anadolu Genel dağılımı: O., B. ve G. Avrupa, K., D. ve G. Afrika, İran, B. Suriye, Kafkasya Bulunduğu kareler: A1 A2 A3 A4 A5 A6 A7 A8 A9 B1 B2 B7 B8 B9 C2 C3 C4 C5 C6

S. columbaria L., Sp. P1. 99 (1753). Figure 18, p. 605. Perennial (sometimes biennial?), stems glabrous or pubescent, often pilose below, reflexed-pubescent above, 30-80 cm. Leaves of sterile shoots, and lower leaves petiolate, lanceolate, oblong or obovate, crenate, dentate or lyrate, rarely entire, glabrous, sparsely or densely pubescent to pannose; upper leaves sessile, 1-2-pinnatisect, segments ovate to linear, entire or dentate. Involucral bracts 8-11, linear, usually shorter than flowers. Capitula 1-5-3 cm diam., ± radiant; flowers lilac, pink, pale yellow or cream, occasionally white, outer flowers with entire petals. Fruiting head spherical or ovoid. Involucel 2-3·5 mm, 8-grooved with hairy ribs; corona c. 1-5 mm with 16-22 nerves. Calyx shortly stipitate; setae 3·5-9 mm long. 1. Plant glabrous subsp. paphlagonica 1. Plant hairy 2. Flowers pink or lilac subsp. columbaria 3. Lower leaves pubescent var. columbaria 3. Lower leaves pannose var. intermedia 2. Flowers yellow or cream subsp. ochroleuca 4. Lower leaves pubescent var. ochroleuca 4. Lower leaves pannose var. webbiana subsp. columbaria. var. columbaria. Syn: S. patens Jordan, Pug. 94 (1852). Ic: Reichb., Ic. Fl. Germ. 12: t. 693 (1850) as Asterocephalus columbarius (L.) Sprengel. Ft. 6-9. Roadsides, wasteland, rocky slopes, s.1.-2600 m. Described from Europe (Hb. Linn. 120/17, photo!). Mainly N. Anatolia. A1(A) Balıkesir: Marmara adasi, A. Baytop 13621! A2(A) Istanbul: Little Alem Da., Hb. Post! Kocaeli: 30 km from Üsküdar to Izmit, c. 50 m, McNeill 218! A5 Samsun: beach nr. Kirazlik, 5 m, Tobey 275! A7 Trabzon: Zigana Da., 2300 m, T, Baytop 14263! A8 Çoruh: Borçka, Tong 458! Rize: mountain forest clearing, 2000 m, Watson 2329! A9 Kars: Yalnizçam Da., above Yalnizçam, 2100-2300 m, D. 32493! B7 Erzurum: N.W. side of Tayran Da., 1500-1800m, Watsonetal. 3257!B8Erzurum:KarakayaDa., 15kmN. of Hinis, 2200-2600 m, Watson et al. 3681! C, W. & parts of S. Europe, N., E. & S. Africa, Iran, W. Syria, Caucasia. var. intermedia (Post) Matthews, comb. nov. Syn: S. ochroleuca L. var. intermedia Post in J. Linn. Soc. (Bot.) 24:433 (1888); S. intermedia (Post) Bornm. in Notizbl. Bot. Gart. Berlin 7:18 (1917). Fl. 8-9. Screes, scrub, 200-2300 m. Type: in rupestribus Amani, G.E. Post. N. Anatolia & Amanus. A6 Tokat: Niksar to Karakus, 1100 m, D. 24929! A7 Gümüşane: Gümüşane, 1220 m, Balls 1619a! Trabzon: Zigana Da., 2300 m, T. Baytop 14300! nr. Trabzon, 200 m, Görz 220. C5 Hatay: Batiayaz to Musa Da., 700 m, Hub.-Mor. 12690. C6 Adana: d. Bahçe, Dildil Da., (Düldül Da.), above Atlik Y., D. 16428! Hatay: Küsliji Da., 750-1520 m, Haradj. 2454! subsp. ochroleuca (L.) Ćelak., Prodr. FL Böhmen. 269 (1873). var. ochroleuca (L.) Coulter, Mem. Dips. 38 (1823). Syn: S. ochroleuca L., Sp. Pl. 101 (1753); S. tenuifolia Roth, Tent. Fl. Germ. 1:59 (1788); S. flavescens Gris. & Schenk in Arch. Naturgesch. 18(l):350 (1852). Ic: Reichb., Ic. Fl. Germ. 12: t. 694 (1850) as Asterocephalus ochroleucus (L.) Sprengel. Fl. 5-9. Fields, rocky slopes, 200-2300 m. Described from Germany. Mainly N. Turkey, Inner & S. Anatolia. A2(E) Istanbul: Chichli (Şişli), hills above Kiathané, 25 vi 1917, B. Post! A2(A) Istanbul: hillsides above Yakacik, 14 ix 1937, B. Post! A3 Bolu: Abant G., 26 vii 1941, B. Post! A4 Zonguldak: Keltepe above Karabük, 1800 m, D. 38853! A5 Samsun: between Vesirköpru and Çakiralan, 690 m, M. Zohary & Orshan 107442! A6 Amasya: Akdağ above Zefe Köy, 1100 m, Tobey 1192! A7 Giresun: 3-4 km S. of Giresun, 200-300 m, Murray 902! A8 Erzurum: pass between Tortum and Oltu, 2300 m, D. 47605! A9 Kars: Ardahan to Hoçuvan, 1800 m, D. 32592! Bl Balikesir: Kaz Da., Quézel et al. B7 Erzincan: 38 km N.W. of Erzincan, 2000 m, Hub.-Mor. 15683. B9 Bitlis: 4 km W. of Reşadiye, 1800 m, D. 46053! C4 Konya: d. Ermenek, Hamitseydi boğaz to Beşköyu, D. 16253! C5 Niğde: nr. Maden, Darrah 364! C6 Adana: Amanus mountains, Hasan Beyli, 1220-1525 m, Haradj. 2279! C. & S. Europe, W. Russia, N. Africa, Lebanon, W. Syria, Siberia, Caucasus. var. webbiana (Don) Matthews, comb. nov. Syn: S. webbiana Don in Bot. Reg. 9: t. 717 (1823); S. ochroleuca L. var. webbiana (Don) Boiss., Fl. Or. 3:132 (1875). Ic: Don, op. cit. (1823). Fl. (4-)5-9. Rocky slopes, open scrub, 100-2500 m. Type: [Turkey Bl Balikesir] Phrygia, Mt. Ida (Kaz Da.), Webb. Turkey-in-Europe & Outer Anatolia. A2(E) Istanbul: Kiathane, 30 vi 1906, Azn.! A2(A) Bursa: 28 km E. of Bursa, 500 m, Sorger 68-50-7! A3 Sakarya: Doğançay, 100 m, D. 36317! A5 Samsun: Kizilirmak after Asarköy, 250 m, Tobey 2233! A6 Ordu: Fatsa to Aybasti, 600 m, Tobey 1364! Bl Balikesir: Mt. Ida (Kaz Da.), Sint. 1883:417! B2 Kütahya: d. Gediz, Şaphane Da., 2000 m, D. 18451! C2 Denizli: Baba Da., 1900 m, D. 18407! C3 Isparta: Dedegöl Da., 1500 m, Sorger 65-43-128! C5 Niğde: Kizil Tepe, 2500 m, Siehe 1895:245! C6 Hatay: Amanus, Köstelli, Akman 3573. Greece, Bulgaria, Yugoslavia. subsp. paphlagonica (Bornm.) Matthews, comb. et stat. nov. Syn: 5. paphlagonica Bornm. in Feddes Rep. Beih. 89:311 (1944). Fl. 7-8. Open woodland, roadsides, 1400-2200 m. Type: [Turkey A4/5 Kastamonu] Paphlagonia: in herbidis montis llkas-dagh (Ilgaz Da.) regionis alpinae abietinae, 2200 m, 14 vii 1890, Bornmüller 2384 (G!). A4 Kastamonu: Ilgaz to Dağları, 1400 m, Khan et al. 668! N. side of Ilgaz Da., 1900 m, D. 38343! Endemic. Euxine (mt.) element. The flowers of subsp. paphlagonica are yellow, as in subsp. ochroleuca. The specimens seen exhibit a remarkably uniform facies and possess constantly elongate-elliptic to lyrate lower leaves. The upper leaves are pinnatisect with linear (sometimes dentate) segments. S. bipinnata C. Koch from Caucasia, is very close to S. columbaria subsp. ochroleuca, and is recorded from E. Anatolia in F1. URSS 24:77 (i'957). I have seen no material, but feel that it is likely that the two are synonymous. A Koch record from Istanbul is probably a misidentification for S. columbaria subsp. ochroleuca.