Asteraceae

Senecio integrifolius subsp. karsianus

Senecio integrifolius subsp. karsianus

Senecio L. Senecio integrifolius (L.) CLAIRV. subsp. karsianus MATTHEWS Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 6 Son çiçeklenme zamanı: 7 Habitat: sulak çayır, su kenarı Minimum yükseklik: 1900 Maksimum yükseklik: 2200 Endemik: endemik Element: Karadeniz (Dağ) Türkiye dağılımı: KD. Anadolu Genel dağılımı: Türkiye Bulunduğu kareler: A2 A3 A4 A5 A7 A8 A9

S. integrifolius (L.) Clairv., Man. Herb. Suisse 241 (1811). Syn: Othonna integrifolia L., Sp. Pl. 925 (1753). Erect perennial with ridged densely or sparsely arachnoid stems 15-80 cm. All leaves arachnoid below, often glabrescent above; basal adpressed to ground, persistent at anthesis, ovate to elliptic, acute or obtuse, base gradually or suddenly attenuate into winged petiole, margin subentire to dentate or crenate; lower cauline oblong-lanceolate, sessile; upper cauline linear to lanceolate, sessile. Inflorescence corymbose, (l-)3-17-capitulate, peduncles 0.5-9 cm. Capitula radiate. Phyllaries 5-10 mm, linear, green or purplish, glabrous or sparsely tomentose. Calyculus absent. Ligules 6-18 mm, yellow or orange, some-times reflexed on flowering. Achenes glabrous or pubescent. 1. Stems densely arachnoid; achenes glabrous; ligules 10-16(-18) 2. Peduncles 0.5-2 cm; phyllaries purplish; ligules orange subsp. aurantiacus var. leiocarpus 2. Peduncles 1.5-9 cm; phyllaries green; ligules yellow subsp. aucheri 1. Stems sparsely arachnoid; achenes pubescent; ligules 17-20 subsp. karsianus subsp. aurantiacus (Hoppe ex Willd.) Briq. & Cavill. var. leiocarpus Boiss., Fl. Or. 3:412 (1875). Syn: S. caucasigenus Schischkin in Not. Syst. (Leningrad) 16:427 (1954). Map 17, p. 161. Fl. 6-7. Grassy slopes, screes, 2135-3350 m. Syntypes: [Turkey A8 Rize] in monte Bousdouandagh Ponti Lazici, 2440 m, Balansa; [Caucasus] in montibus Artschikala et Saridagh Daghestaniae australis, 3200-3350 m, Ruprecht; monte Elbrus, Radde; Kasbek, 3050 m, Rehm., forma magis cespitosa 4-6-pollicaris. N.E. Anatolia. A7 Giresun: Balaban Da. (Kilinç Tepe) above Tamdere, 2900 m, D. 20540! A8 Gümüşane: Kara Kaya Da., N. of Bayburt, 3350 m, Balls 1815! Rize: Rize to Ispir, 3000 m, Stn. & Hend. 6265! Erzurum: Mesçit Da., S.E. of Ispir, 2900-3000 m, Barclay 924! Çoruh: Kum Kurun Da. nr Melo above Artvin, 2135 m, Apold et al. 113! A9 Çoruh/Kars: pass between Ardanuç and Ardahan, 2400 m, Kerck 40/7! Caucasia. Euxine element. subsp. aucheri (DC.) Matthews in Notes R.B.G. Edinb. 33:434 (1975). Syn: S. aucheri DC, Prodr. 6:361 (1838). Map 17 p. 161. Fl. 6-7. Rocky slopes, meadow, woodland, 1800-2200 m. Type: [Turkey A2(A) Bursa] in Olympo Bithyniae (Ulu Da.), Aucher (K!). N. Anatolia, W. of 35°E. A2(A) Bursa: Ulu Da., Moore 7290! ibid., Pichler! ibid., Sorger 62-95-2! A3 Bolu: Ala Da., Kartal Kaya Tepe, 2100-2200 m, D. 37361! A4 Çankiri: Küçük Ilgaz Da., 1800-2000 m, Schiechtl {Hub.-Mor. 17304)! Kastamonu: N. side of Ilgaz Da., 2100 m, D. 38452.! A5 Kastamonu: Ilgaz Da., 2000 m, Darrah 7! Greece. Euro-Sib. (Euxine?) element. subsp. karsianus Matthews in Notes R.B.G. Edinb. 33:257 (1974). Map 17, p. 161. Fl. 6-7. Water meadows, stream-sides, 1900-2200 m. Type: Turkey A9 Kars: Maçuvan between Kars and Ardahan, 1900 m, edge of water meadow, 30 vi 1957, Davis & Hedge, D. 30474 (holo. E!). N.E. Anatolia. A9 Kars: 7 km from Sarikamiş to Karaurgan, 2200 m, D. 46617! Kisir Da., above Susuz, 2000 m, D. 30522! Endemic. Euxine (mt.) element. Three specimens from A4 Kastamonu: Ilgaz Da. belong to S. integrifolius but are intermediate between subspp. aurctntiacus and aucheri. They have yellow ligules but phyllaries which are purplish (Markgraf s.n.!) or a mixture of green and purple (Edmondson 514! 519!). A specimen from A8 Erzurum: nr Ispir, Huet, was assigned to var. aurantiacus by Boissier (Fl. Or. 3:412, 1875). Subsp. aurantiacus sensu stricto has hairy achenes and occurs in the Eastern Alps and Carpathians; the Turkish specimen was presumably misidentified. A record of S. campestris (Retz.) DC. from A3 Bolu (Ala Da., Wiedemann) is most probably referable to subsp. aucheri. S. integrifolius is a very variable species which has been much subdivided. Subsp. integrifolius is found in most of C. & E. Europe. The related European species form a taxonomically very difficult group and further study of the whole complex is needed.