Lamiaceae

Teucrium chamaedrys subsp. tauricolum

Teucrium L. Teucrium chamaedrys L. subsp. tauricolum RECH. FIL. Ömür: Çok yıllık Yapı: ot Hayat formu: İlk çiçeklenme zamanı: 7 Son çiçeklenme zamanı: 8 Habitat: kuru kayalık yamaçlar, uçurumlar ve hareketli kayalıklar Minimum yükseklik: 500 Maksimum yükseklik: 3000 Endemik: endemik Element: D. Akdeniz Türkiye dağılımı: GB. ve G. Anadolu Genel dağılımı: Türkiye Bulunduğu kareler :A1 A2 A3 A4 A5 A6 A7 A8 A9 B1 B2 B3 B4 B5 B6 B7 B8 B9 C1 C2 C3 C4 C5 C7 C8 C9

T. chamaedrys L., Sp. Pl. 565 (1753). Rech. fil. in Bot. Archiv 42:338-389 (1941). Syn: T. officinale Lam., Fl. Fr. 2:414 (1779); T. pseudochamae-drys Wender. in Flora 9:358 (1826). Ic: Bot. Archiv 42:417-420 (1941). Suffrutescent perennial herb, often rhizomatous, 5-50 cm, with very variable indumentum. Leaves oblong or obovate-oblong, cuneate at base, usually crenate-dentate or shortly lobulate, lobes entire or toothed. Verticillasters (2-)4-8-flowered, in dense or ± lax terminal racemes, floral leaves (bracts) usually distinct from cauline leaves, entire or toothed; pedicels equalling orusually shorter than calyx. Calyx tubular-campanulate, subgibbous at base, teeth lanceolate-triangular, ± subulate-tipped, margins usually densely short-haired and sparsely long-hairy, or only long-hairy, or glabrous, usually purplish. Corolla reddish-purple, 2 x calyx. Stamens at least as long as upper lateral lobes 1. Bracts all overtopping calyces 2. Leaves subglabrous or shortly hispid beneath subsp. lydium 2. Plant bearing long eglandular or glandular hairs 3. Calyx teeth glabrous or sparsely long-haired subsp. trapezunticum 3. Calyx teeth densely covered with short and long hairs subsp. chamaedrys 1. Bracts mostly shorter than calyces 4. Plant shortly retrorse- and crispate-hairy; stem hairs diminishing in length downwards subsp. sinuatum 4. Indumentum not as above 5. Upper bracts entire, all ciliate-margined; stem hairs not diminishing in length upwards;leaves ± discolorous subsp. syspirense 5. Bracts all dentate, not usually ciliate-margined; stem hairs diminishing in length upwards;leaves concolorous subsp. tauricolum subsp. chamaedrys. Syn: T. chamaedrys L. subsp. euchamaedrys Rech. fil. in Bot. Archiv 42:344 (1941); T. chamaedrys L. var. ramigera Wierzb. in Čelak. in Bot. Centr. 14:220 (1883); T. chamaedrys L. subsp. parviflorum Formánek in Verh. Nat. Ver. Brünn. 34:325 (1895), incl. var. viridescens Formánek, op. cit. 36:84 (1897); T. chamaedrys L. var. glanduliferum Hausskn. in Mitt. Thür. Bot. Ver. N.F. 11:31 (1897). Fl. 6-9. Open forests, cliffs, slopes, steppe, s.I.-1600 m. Described from Germany, Switzerland and France (Hb. Linn. 722/19!). N. Turkey, C. & S. Anatolia. A1(E) Kırklareli: 15 km W. of Iğneada, Bauer et al. 1968: 2436(!). A2(E) Istanbul: Baltalimani, 1936, B. Post! A2(A) Istanbul: Göztepe, Körfez, 1939, B. Post! Kocaeli: Gök Da., Kel Tepe, 1500-1600 m, Dingier 239 (var. glanduliferum). A3 Bolu: Akyarma, 1560 m, Uot-ila 20004! A4 Çankiri: Çerkeş, Kilinç 222! A5 Kastamonu: Tosya, 1600 m, Khan et al. 649! A6 Samsun: Borabay, 900 m, Tobey 2309! Samsun, 50 m, Hand.-Mazz. 130 (var. glanduliferum). Bl Izmir: Boz Da., 1962, Regel! B2 Kütahya: above Simav, 1300 m, E. Anglia Exped. E9! B3 Konya: Akşehir, Sultan Da., 1450-1500 m, Hub.-Mor. 10154! B4 Ankara: 15 km from Ankara to Polatlı, Dudley (D. 35999a)! B5 Nevşehir: Nar, 3 km N. of Nevşehir, 1250 m, Roper 80! Cl Aydin: Paşa Y., 1020 m, Uslu 4294! C2 Denizli: Denizli to Acipayam, Dudley (D. 35336)! C3 Antalya: Akseki to Manavgat, 590 m, Hub.-Mor. 10153! C4 Konya: Bozkir, 1100 m, Hub.-Mor. 10152! C5 Adana: 5 km N. of Pozanti, 1200 m, Aberdeen Univ. Amanus Exped. Al 34! Most of Europe except N. Euro-Sib. element. A very variable subspecies, glandular plants (var. glanduliferum) being less common in Turkey than the typical eglandular forms. Subsp. chamaedrys is easily confused with subsp. tauricolum, from which it differs by its ciliate and usually longer bracts, the whole plant having longer indumentum. subsp. lydium O. Schwarz in Feddes Rep. 36:132 (1934). Ic: Rech. fil., op. cit. t. 1 f. 5, t. 3 f. 42 (1941). Fl. 6-7. Pinus nigra forest, macchie, stony serpentine slopes, open slopes, 500-1700 m.Syntypes: [Turkey Bl Izmir] Lydia, Nifdag, in pascuis pinetigeris solo calcareo et schistoso, (600-) 1000-1400 m, [7 vii 1933], O. Schwarz 818 (JE?); Çiplakdag, in pratis herbosis solo schistoso, 1400-1600 m, O. Schwarz 934 (JE?). N.W. Turkey, W. & S.W. Anatolia. A2(E) Istanbul: Aydos Da., Berk & T. Baytop (ISTE 3933)! A2(A) Kocaeli: Gebze to Izocam, Ertem & Özhatay (ISTE 22260)! Bl Çanakkale: mt. Ida (Kaz Da.), Kareikos, Sint. 1883:586! Izmir: Kemalpaşa, Alava 4951! B2 Kütahya: Murat Da., 1500 m, M. & D. Zohary 5523(!). C2 Antalya: Kizilca Da., Küçüklu Çayi valley, 1500 m, McPhail & Watson 5706(!). C3 Antalya: Manavgat to Akseki, 500 m, Dudley (D. 35766)! Bucak to Hafizpaşa, Hub.-Mor. 5858! Isparta: W. side of Beyşehir G., 1200 m, Sorger 6541-1! Greece (Samothrace). E. Medit. element. Very closely related to subsp. chamaedrys, but stem with spreading and subdeflexed hairs (spreading hairs usually longer than stem diameter) and inflorescence more congested; leaves hispidulous or subglabrous beneath. subsp. trapezunticum Rech. fil. in Bot. Archiv 42:369, t. 1 f. 9, t. 3 f. 46 (1941). Fl. 6-8. Rocky slopes and banks, s.l-1500 m. Type: [Turkey A7 Trabzon] Trapezunt (Trabzon), Matardjik, Sintenis 1384 (iso. B, K!W, WU). Mainly N.E. Anatolia. A5 Sinop: Boyabat to Sinop, 1150 m, D. 38069 (glandular)! A6 Ordu: Fatsa to Aybasti, 350 m, Tobey 1325! A7 Trabzon: Karakor, 350 m, Balls 473 (glandular)! A8 Trabzon: Sürmene to Of, Hub.-Mor. 16355! Çaykara, Soğanli Da., 1300 m, D. 32086 (glandular)! Rize: 1 km N. of Ikizdere, 500 m, Hub.-Mor. 16356! Çoruh: Artvin, 600 m, D. 30010! Caucasia. Euxine element. Although originally described as eglandular, glandular variants of this subspecies are frequent. Subsp. trapezunticum is certainly very closely related to the Caucasian subsp. nuchense (C. Koch) Rech. fil. (Bot. Archiv 42:370, 1941), from which it may be distinguished by its long lax stem indumentum and often bifid upper bracts. Grossheim (Fl. Kavk. 7: map 342) maps only subsp. nuchense from Çoruh province, but all material seen from there is referable to subsp. trapezunticum as delimited by Rechinger. Intermediates between these two subspecies have been seen from A7 Giresun (Tamdere to Yavuzkemal, nr Karinca, 1500 m, D. 20731!) and between the same two subspecies and subsp. syspirense from A8 Rize (Ikizdere, 2100 m, D. 20998!). subsp. tauricolum Rech. fil. in Bot. Archiv 42:376, t. 1 f. 13, t. 3 f. 50 (1941). Syn: T. chamaedrys sensu Boiss., Fl. Or. 4:816 (1879) p.p. non L. (1753) & var. canum Boiss., loc. cit. (1879) p.p. non T. canum Fisch. & Mey. (1835); T. syspirense sensu Bornm. in Mitt. Thür. Bot. Ver., N.F. 37 :7 (1927) p.p. non C. Koch (1848); T. chamaedrys L. subsp. tauricolum Rech. fil. var. glaberrimum Rech. fil., op. cit. 377 (1941)! Fl. 7-8. Dry rocky slopes, cliffs and screes, 500-2300 m. Type: [Turkey C5 Içel] alpes Bulgardagh, in apricis collinis ad pagum Güllek frequens, 1060 m, Kotschy 77a (holo. W). Mainly S.W. & S. Anatolia. B3 Afyon: Çay, Sultan Da., Yalt. (ISTO 1068)! C2 Denizli: Babadağ, 2200 m, D. 18394! Antalya: Elmali, Bourgeau 214! C3 Antalya: Irmassa pass between Manavgat and Beyşehir, 1470-1525 m, 18 viii 1972, Shmida & Luria(!). C4 Içel: 6 km S. of Uzuncaburç, 500 m, Sorger 71- 25-23! C5 Adana: Pozantı, 850 m, Aberdeen Univ. Amanus Exped. B2 103! Içel: Bolkar Da., 1 km W. of Arslanköy, 1600 m, Peat et al. 29-1-76(!). Niğde: N. side of Taurus, Madenköy, 1670 m, Spitzenberger 1970:136 (!). Hatay: Amanus, above Arsuz, 610 m, Kotschy 1862:149 (type of var. glaberrimum). C6 Maraş: Akher (Ahir) Da., 1830-2135 m, Haradj. 1708! Adana: Saimbeyli, 1100 m, D. 19887 (var. glaberrimum Rech. fil.)! Hatay: Soğuk Oluk to Iskenderun, Amanus Da., Akman 3381! Amanus, Belen, 500-520 m, Hub.-Mor. 12760 (var. glaberrimum)! C7 Adiyaman: Nemrut Da., nr Kahta, 1600-2500 m, Hand.-Mazz. 2103. Endemic. E. Medit. element. Var. glaberrimum, with glabrous leaves and very short crispate stem indumentum, is apparently confined to the Amanus and Anti-Taurus. subsp. syspirense (C. Koch) Rech. fil. in Ann. Naturh. Mus. Wien 51:427 (1941). Syn: T. syspirense C. Koch in Linnaea 21:704 (1848); T. chamae-drys L. var. canum Boiss., Fl. Or. 4:816 (1879) p.p. non T. canum Fisch. & Mey. (1835); T. chamaedrys L. var. hirsuta Čelak. in Bot. Centr. 14:220 (1883); T. chamaedrys L. var. gymnobotrys Celak. op. cit. 221 (1883)! Fl. 6-8. Limestone or igneous rocky slopes, steppe, 90-2430 m. Type: [Turkey A8 Erzurum] Horasan, im Gaue Sber (Ispir), sehr haufig auf porphyr, 915-2430 m, [1844], C. Koch. Inner & adjacent N. & S. Anatolia. A4 Kastamonu: 5 km W. of Kastamonu, 900 m, D. 38771! A5 Sinop: Gerze, 30 km S. of Sinop, Hub.-Mor. 16358! A6 Amasya: Erbaa to Koçak, 300 m, Tobey 2177! A7 Gümüşane: Gümüşane, Sint. 1894:6099! A8 Erzurum: 20 km N. of Aşkale, 1900 m, Kukkonen 8053! B2 Kütahya: 18 km S. of Kütahya, 1000 m, D. 36935! B3 Afyon: Sultan Da., Bornm. 1899:5422. B4 Ankara: Beynam forest, D. 13013! B5 Kayseri: Talas, 1230-1270 m, Hub.-Mor. 10794! B6 Maraş: 5 km W. of Elbistan, 1070 m, Hub.-Mor. 12579! B7 Malatya: 3 km E. of Malatya, 950 m, Hub.-Mor. 8956! Sivas: Divrik (Divriği), c. 1000 m, Bornm. 1893: 3516 (type of T. ixodes, photo!). B8 Erzurum: Palandöken Da., 2300-2350 m, D. 47352! B9 Van: Toprakkale, 1620-1800 m, Hub.-Mor. 8958! C2 Antalya: d. Elmali, Bey Da., Çamçukuru Y., 1750 m, Khan et al. 324! C3 Burdur: 5 km W. of Burdur, 900 m, Hub.-Mor. 8402! C4 Konya: Sara nr Ermenek, 1872, Péronin 186 (type of var. gymnobotrys)! C7 Malatya: 74 km S.W. of Malatya, 1550 m, Sorger 71-45-27! Transcaucasia, N. Iran & Khorassan. Ir.-Tur. element. This rather distinctive subspecies differs from the others in its discolorous leaves, and usually entire bracts which (except the lowest pair) are equal to or shorter than the flowers; T. ixodes Bornm. is only a glandular variant. It is possible to see many intermediate forms between this subspecies and subspp. tauricolum, chamaedrys and trapezunticum, mainly in contact areas; they are listed at the end of the species. subsp. sinuatum (Čelak.) Rech. fil. in Bot. Archiv. 42:378 (1941). Syn: T. sinuatum Čelak. in Bot. Centr. 14:218 (1883); Boiss., Fl. Or. Suppl. 363 (1888); T. divaricatum sensu Rech. fil. in Ann. Naturh. Mus. Wien 49:278 (1939) non Sieber (1821). Fl. 6-8. Igneous and limestone slopes, steppe, road and field sides, 1150-2600 m. Type: [N. Iraq] in rupestribus mont. Gara Kurdistaniae, 2 viii 1841, Kotschy [351] (holo. W). E. Anatolia. A8 Erzurum: nr Ispir, 1200 m, Barclay 797! A9 Kars: Karakurt to Sarikamiş, 1780 m, D. 46483! B6 Sivas: Gürün, 1250-1300 m, D. 21955! B7 Tunceli: Munzur Da., 2000 m, D. 31302! B8 Muş: Karduchia (Kurtik) Da., 1375 m, Kotschy 1859: Suppl. 692. B9 Ağri: E. of Tahir pass, 2150 m, Lamond 2574! C8 Sürt: Siirt, 1150 m, Frödin 1939:203. C9 Hakkari: Çukurca, 1200 m, D. 44747! N. Iraq. Ir.-Tur. element. Very closely related to subsp. syspirense, but indumentum shorter and intemodes usually longer than verticillasters.