| Asteraceae |
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Anthemis cretica L. subsp. anatolica (BOISS.) GRIERSON |
 Anthemis cretica L. subsp. anatolica (BOISS.) GRIERSON |
Anthemis L. |
| A. cretica L., Sp. Pl. 895 (1753) non (L.) Nyman (1854-55). Syn: A. montana L., Sp. PL ed. 2:1261 (1763), nom. illegit. A. orientalis auct. non (L.) Degen, Degen It. Turc. (Exsicc, 1890) non Anacyclus orientalis L. Shortly rhizomatous perennial herb, sometimes woody at base, subglabrous or ± densely greyish- or white-pubescent. Stems decumbent, ascending or erect, 10-30(-40) cm, simple or branching. Leaves pinnatisect, up to 5 cm, primary segments usually 3-paired, simple or more often divided into 3-5(-9) oblanceolate ± obtuse secondary segments. Capitula radiate or discoid. Involucre generally 0.75-1.5 cm broad; phyllaries ovate or lanceolate, ± acute, margins ± scarious, colourless, pale or dark brown. Receptacle often becoming somewhat conical at maturity; paleae lanceolate, carinate. Ray flowers when present white, usually fertile; ligules elliptic, 0.5-1.25 cm. Disc flowers yellow, corolla-tube usually becoming inflated at maturity. Achenes obpyramidal, ± squarish in section, 1.5-3.5 mm, smooth, sulcate or especially the outer subtuberculate. Pappus auriculate or coroniform, ± absent or up to 1 mm, scarious. Type: Hb. Cliff. 415/2! This is a highly polymorphic species in which a number of different taxa are recognizable and have been variously treated at specific or infraspecific levels by previous authors. The following is a tentative classification but, such is the degree of intergradation, no satisfactory scheme seems possible without some biosystematic study; numerous intermediates occur between the subspecies recognized here; radiate and discoid variants are treated informally and epithets have not been given to them as it has seemed difficult to avoid confusion with those applied, perhaps slightly differently, in Europe. The external distribution has been curtailed due to lack of information. The illegitimacy of the name A. montana L. (previously applied to this widespread species complex) is discussed by Grierson in Notes R.B.G. Edinb. 33: 212 (1974). 1. Leaf segments linear-lanceolate, c. 0.5 mm broad, acute or acuminate (some-times broader but then confined to islands); radiate and discoid plants e. subsp. leucanthemoides 1. Leaf segments oblanceolate or obovate, usually more than 0.5 mm broad, obtuse or subacute 2. Lower or median leaves usually more than 2 cm 3. Phyllaries with dark brown or blackish margins; plants radiate and discoid, or radiate only 4. Taller plants (20-)30-40 cm tall; radiate and discoid plants 5. Leaf segments 1 .5 mm or less broad b. subsp. albida 5. Leaf segments (l-)2-3.5 mm broad g. subsp. cassia 4. Smaller plants 15(-20) cm tall (or less from higher alpine habitats); radiate plants only 6. Leaves usually glabrous (Ulu Da.) j. subsp. carpatica 6. Leaves ± densely whitish pubescent (N.E. Anatolia) k. subsp. iberica 3. Phyllaries with whitish or pale brown margins; plants radiate and discoid 7. Primary leaf segments divided into 7 or more segments h. subsp. absinthifolia 7. Primary leaf segments divided into 3 or 5 segments 8. Stems elongate, 30-45 cm, much branched i. subsp. candicans 8. Stems 10-20(-40) cm, simple or branched only near base 9. Plants usually with several leafy sterile shoots; leaf segments ± contiguous a. subsp. pontica 9. Plants without or with few sterile shoots; lear sentents ± distant f. subsp. umbilicata 2. Lower and median leaves usually less than 2 cm 10. Phyllaries with whitish or pale brown margins; radiate or discoid plants 11. Indumentum dense, leaves often greyish or white; marginal achenes usually subtuberculate d. subsp. anatolica 11. Indumentum ± sparse, leaves green; marginal achenes smooth c. subsp. tenuiloba 10. Phyllaries with dark brown or blackish margins; radiate plants only subsp. argaea d. subsp. anatolica (Boiss.) Grierson in Notes R.B.G. Edinb. 33:428 (1975). Syn: A. anatolica Boiss., Diagn. ser. 1(11): 10 (1849)! A. anatolica Boiss. var. discoidea Boiss., Fl. Or. 3:295 (1875) (non A. montana var. anatolica Boiss.). Figure 9, p. 181. Small herbs often greyish-pubescent, sometimes quite strongly woody at base. Stems decumbent or ascending, 10-20(-30) cm. Leaves generally less than 2 cm, segments sometimes entire, but generally temately divided, sometimes ultimate segments congested. Capitula radiate or (in type) discoid. Involucre 0.7-1 cm broad; phyllaries pale or with narrow brownish margins. Achenes, especially outer ones, ± tuberculate; corona 0.25-0.5 mm. Fl. 5-7. Steppe, roadsides and limestone hillsides, 280-2285 m. Syntypes: [Turkey C2 Denizli] circa Gheyra, Boissier (G!); [B1 Manisa/ Izmir] in monte Sipylo (Manisa Da.), Balansa 24 (G!) Radiate variants. C. & S.W. Anatolia. B4 Kirşehir: 23 km S.E. of Şerefli Koçhisar, 900 m, Ehrendorfer 62-l/77a-7! C2 Burdur: 81 km from Fethiye to Dirmil, 1100 m, Hub.-Mor. 5430 (approaching subsp. leucanthemoides)! Antalya: Elmali, 1860, Bal. 166! Muğla: between Aydin and Muğla, 10 km from Yatağan, 280 m, Hub.-Mor. 12375! Denizli: Cadmus (Honaz Da.), vi 1842, Boiss. (type of A. anatolica)! C3 Antalya: 29 km S.W. of Korkuteli, Sorger 65-23-7! C4 Konya: 38 km W. of Konya, 1500 m, Sorger 64-36-11! C5 Niğde: Ala Da., 1500-2060 m, Parry 14! Discoid variants. Mainly C, W. & S. Anatolia. B2 Manisa: 24 km from :Inegöl to Buldan, Hub.-Mor. 5390! B4 Konya: Cihanbeyli, D. 18641! B5 Nevşehir: 12 km from Gülşehir (Arapsun) to Nevşehir, 500 m, McNeill 389! B6 Sivas: Gürün, 1400 m, Stn. & Hend. 5257! B7 Malatya: 15 km N.E. of Malatya, 1100 m, M.Zohary 37-2130! C2 Burdur: Kizilhisar to Yeşilova, 9 km W. of Salda G., 1180 m, Hub.-Mor. 17292! Isparta: Eğridir, Barla Da., 2250 m, Khan et al. 369! Muğla: d. Fethiye, Baba Da., D. 13664! C3 Antalya: Tahtali Da., 2100 m, D. 14145! C4 Antalya: Geyik Da., 2285 m, D. 14528! C7 Malatya: 74 km S.W. of Malatya, 1550 m, Sorger 71-45-5! |