Asteraceae |
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Carduus onopordioides FISCH. EX BIEB. subsp. onopordioides FISCH. EX BIEB. |
Carduus onopordioides FISCH. EX BIEB. subsp. onopordioides FISCH. EX BIEB. |
Carduus L. |
C. onopordioides Fisch. ex Bieb., Fl. Taur.-Cauc. 3:552 (1819). Stout biennial (?), 20-50(-90) cm, subglabrous.' Stem conspicuously winged with triangular-lanceolate, stoutly spined lobes, sparsely paniculate above. Cauline leaves pinnatifid to pinnatipartite, thick, primary segments distant (separated by squarish sinuses), triangular and lobed, stoutly spined (spines 3-7 mm). Peduncles up to 5 cm, strongly spiny-winged nearly up to subsolitary capitula; lateral capitula subsessile. Involucre subglobose to broadly campanu-late, 18-32 x 18-50 mm (incl. spines), gradually imbricate, weakly floccose or glabrous, outer and median phyllaries variable, lanceolate-subulate, often carinate, long-spined and falcately patent or recurved, or lanceolate and shortly spined, ± patent; inner scarious, purple, linear-lanceolate or lanceolate. Corollas zygomorphic, purplish-pink, 15-19 mm, shorter to longer than median phyllaries. Achenes (subsp. onopordioides) 5 mm; pappus c. 16 mm. 1. Involucre 25-32 cm (incl. spines); median phyllaries lanceolate-subulate, long-spined, falcately patent, as long as or longer than flowers; inner phyllaries linear-lanceolate, long-attenuate, about as long as flowers subsp.onopordioides 1. Involucre 18-20 cm (incl. spines); median phyllaries lanceolate, less prominently nerved, more abruptly narrowed into a shorter weaker ± patent 1-3 mm spine; inner phyllaries lanceolate and more abruptly tapered, longer than median phyllaries but much shorter than flowers subsp. turcicus subsp. onopordioides. Icr:Fl. URSS 28: t. 1 f. 3 (1963); Fl. Gruzii 8: t. 411 (1952). Map 45, p. 433. Alt. c. 2100 m. Type: in Iberia, Wilhelms (LE, G). E. Anatolia, rare. B9 Van: 2 km E. of Hoşap, stony gully in steppe, 2100 m, D. 44566! Transcaucasia, N.W. Iran. Ir.-Tur. element. The Turkish specimen has more slender and arachnoid phyllaries than is usual in Caucasia and Iran. |