Asteraceae

Asteraceae

Centaurea urvillei DC.subsp. nimrodis (BOISS. ET HAUSSKN.) WAGENITZ

Centaurea L.
Centaurea urvillei DC.
subsp. nimrodis (BOISS. ET HAUSSKN.) WAGENITZ
Ömür: Çok yıllık
Yapı: ot
Hayat formu:
İlk çiçeklenme zamanı: 6
Son çiçeklenme zamanı: 6
Habitat: ?
Minimum yükseklik: 1000
Maksimum yükseklik: 2000
Endemik: endemik
Element: İran-Turan
Türkiye dağılımı: GD. Anadolu
Genel dağılımı: Türkiye
Bulunduğu iller
Bulunduğu kareler:A2 A3 A4 A5 A6 B1 B2 B3 B4 B5 B6 B7 B10 C3 C4 C5 C6 C7

C. urvillei DC, Prodr. 6:592 (1838).
Short-lived perennial (or biennial ?), 5-30 cm, stem simple or branched from near base. Leaves slightly arachnoid to distinctly tomentose; lyrate with triangular, rhombic or oblong-ovate terminal segments and few to numerous lanceolate, oblong or lyrate lateral segments, interspersed with small lobes. Involucre 20-40 x 15-40 mm, ovoid to globose. Appendages very variable (see subspecies) with numerous cilia (2-)3-5(-7) mm, ending in 10-30 mm spine (incl. spine). Flowers rose-purple or whitish, marginal slightly radiant. Achenes 4-6 mm; pappus (5-)8-ll(-13) mm. Bristles of the receptacle 15-20 mm. 2n = 20, 40.
A very polymorphic species the treatment of which can be only provisional. I have distinguished five subspecies connected by transitional forms although the extremes (subsp. urvillei and subsp. hayekiana) look very different and could easily be taken for species. The geographical differentiation is only partial, especially where the areas of subsp. urvillei and subsp. armata overlap. C. urvillei is endemic to Turkey and closely related to C. lydia and C. raphanina sensu lato.
1. Appendages (excl. cilia) 2-4(-5) mm broad at base, rarely more than 20 mm
long; stem very short or elongate and mostly rather slender
  2. Lateral segments of leaves entire or dentate; capitula few (1-4) subsp. urvillei
  2. Lateral segments of leaves at least partly lyrate; plants usually with more than 4 capitula subsp. stepposa
1. Appendages 4-9 mm broad at base, usually over 20 mm long; stem short or elongated, always stout
   3. Appendages 4-6(-7) mm broad at base, with (5-)8-12 cilia on each side subsp. armata
   3. Appendages (5-)6-9 mm broad at base, with 11-15 cilia on each side
     4. Appendages abruptly contracted into terminal spine, usually straw-coloured; flowers mostly whitish subsp. hayekiana
     4. Appendages gradually narrowed into spine, brown; flowers rose-purple subsp. nimrodis
subsp. nimrodis (Boiss. & Hausskn.) Wagenitz in Willdenowia 6:494 (1972). Syn: C. nimrodis Boiss. & Hausskn. in Boiss., Fl. Or. 3:664 (1875)! Fl. 6.
Type: [Turkey C7 Urfa] Nimrud Dagh supra Orfa (Urfa), 1865, Haussknecht (holo. G!).
Southern E. Anatolia. C7 Urfa: Nimrud Da. (Nemrut Da.), Sint. 1888:1526!
Ir.-Tur. element. One further gathering (C9 Hakkari: 5 km from Hakkari to Kaval, 1850 m, D. 45424!) differs in several characters and may represent a subspecies of its own. Further study is also necessary for other plants from E. Anatolia. The two following collections represent tall purple-flowered plants with few large capitula. They have been assigned to C. deinacantha Boiss. & Hausskn. from Iraq (perhaps only a subspecies of C. urvillei), but the spines are not so robust as in this species: C7 Adiyaman: Nimrud Da. (Nemrut Da.), below Urik (Horik), 1000-1200 m, Hand.-Mazz. 2168! B1O Bitlis: Bitlis, 1400 m, Frödin 79!