Asteraceae |
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Echinops L. |
E. pungens Trautv., Echin. Gen. 18 (1833). Stems 1-4, usually branched above, ridged, 50-100 cm, glabrous, with glandular hairs, or arachnoid to lanate. Leaves linear-elliptic in outline, up to 40 x 20 cm, usually 1-2-pinnatisect, arachnoid-glandular or -eglandular above, lanate below with glandular hairs on veins; lower leaves petiolate, upper amplexicaul, some-times simple; spines numerous on all parts of leaf and petiole. Heads up to 8 cm, pale blue or whitish, with short or long leafless peduncles. Capitulum c. 2.5 (-3.5) cm; brush ± sparse, less than 1/2 as long as capitulum, 7-10(-16) mm. Phyllaries c. 20(-28), glabrous; outer c. 10(-18) mm; median c. 15(-25) mm, acute; innermost free or scarcely connate at base, 15-20(-33) mm. Corolla paleblue to whitish; tube 7-9 mm with glandular hairs; lobes 9-11 mm. Achenes 8-15 mm. Pappus bristles connate to about 1/3 of their length. Fl. 6-8. Rocky limestone, serpentine and igneous slopes in steppe, fallow fields, roadsides, 1100-2700 m. A common species in eastern Anatolia, variable in size, degree of leaf division, indumentum and phyllary size and shape. Although usually a perennial, biennial forms do occur. Four varieties are recognized, two of which (var. polyacanthus and var. transcaucasicus) were given specific rank in Fl. URSS (27:34, 36, 1962). 1. Stems entirely glabrous below var. polyacanthus 1. Stems glandular, arachnoid-tomentose or lanate 2. Upper cauline leaves simple, lobed or pinnatifid var. transcaucasicus 2. Upper cauline leaves pinnatisect or pinnate 3. Stems green with an indumentum almost entirely of glandular hairs var. adenoclados 3. Stems grey-white with an eglandular or glandular and eglandular indu-mentum var. pungens var. adenoclados Hedge in Notes R.B.G. Edinb. 33:239 (1974). Type: Turkey B9 Bitlis: Kotum, 1800 m, Davis & O.Polunin, D. 24549 (holo. E! iso. K!). S.E. Anatolia. B9 Van: d. Başkale, Ispiriz Da., 2700 m, D. 23723! C10 Van: 20 km S. of Başkale on road to Hakkari, D. 24517! Endemic? Ir.-Tur. element. Closest to var. polyacanthus but stems glandular throughout, without any arachnoid or lanate indumentum. |