Boraginaceae |
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Rindera PALLAS |
R. lanata (Lam.) Bunge in Mem. Sav. Etr. Acad. Petersb. 7:415 (1851). Syn: Mattia umbellata C.Koch in Linnaea 17:302 (1843) non Schultes (1809); M. brachyantha Boiss., Diagn. ser. 1(11): 127 (1849); Rindera pubescens C.Koch in Linnaea 23:648 (1849). Ic: Post, Fİ. Syr. f. 537 (1896). Figure 9, p. 245. Erect perennial herb with stout, tapering taproot. Stems simple or rarely branched in lower part, 15-55 cm, sparsely hairy to glabrous. Basal leaves long-petiolate, ovate, oblong or linear, lamina 20-150x2-25 mm, acute to obtuse, base attenuate into petiole, softly tomentose or pilose, without tuberculate-based trichomes, or glabrescent with numerous calcified tubercles; lower cauline oblong or linear, attenuate at base, upper broadly ovate, acuminate, amplexicaul. Cymes numerous, forming a very large terminal corymb. Pedicels greatly enlarging in fruit. Calyx 3-5-8 mm, lobes ± ovate, very densely white-lanate. Corolla pink, drying deep blue, 7-12 mm, campanulate-cylindrical, tube $-£ χ limb. Faucal appendages subquadrate, apex emarginate. Stamens included, filaments all equal, 2/3-11/2 anther length. Style 7-16 mm, usually exserted. Nutlets (2 often aborting) ± orbicular, 14-26 χ 15-23 mm, smooth, wing with smooth or undulate often blue margin, without glochids. 1. Basal leaves oblong, elliptic or ovate, 25-140 χ 5-25 mm var. lanata 1. Basal leaves linear or linear-lanceolate, 20-130 χ 2-12 mm var. canescens var. canescens (A.DC.) Kusn. in Trav. Mus. Bot. Acad. Petersb. 7:48 (1910), incl. subvar. detonsa (Bornm.) Brand in Engler, Pflanzenreich 78 (IV.252):73 (1921). Syn: Mattia canescens A.DC. in DC, Prodr. 10:168 (1846); M. eriantha Ledeb., Fl. Ross. 3:173 (1867); Rindera tanata (Lam.) Bunge var. eriantha (Ledeb.) Kusn. in Trav. Mus. Acad. Sci. Petersb. 7:47, t. 1 (1910); R. lanata (Lam.) Bunge var. stenophylla Bornm. in Beih. Bot. Centr. 28:473 (1911). Ic: Takht. & Fedorov, Fl. Erevana ed. 2: t. 66 (1972), as Rindera lanata. Fl. 5-8. Banks, meadows, pastures, igneous slopes, 1300-3050 m. Syntypes: 'In Oriente', Michaux; [W. Iran] inter Kermancha et Amadan, Olivier & Bruguiẻre (G-DC, photo!). Mainly S. & Inner Anatolia. A7 Gümüşane: Moaldasdagh (Mağaldas Da.), Sint. 1894:5834 p.p.! A8 Gümüşane/Erzurum: Bayburt to lspir, Bourgeau 196! A9 Kars: Sarikamiş, 2100 m, D. 30796! B4 Ankara: Hüseyingazi, 1200 m, Markgraf 11030! B5 (?)Kayseri: Hisarcik to Tekir Y., 1700-2100 m, Çetik 4181! B7 Erzincan: Kara Da., 1520 m, Balls 1081a! B8 Erzurum: 6 km E. of Tercan, 1829 m, Furse & Synge 43! B9 Van: d. Gevaş, Artos Da., 2900 m, D. 22768! B10 Ağri: pass S. of lğdir, Mathews & Tomlinson 4283! C3 lsparta: Çiçek Da., 1600 m, Sorger 61-5-99! C4 Konya: Serai Dagh (Saray Da.) nr Konya, c. 1600 m, 12 v 1902, Zederbauer! C7 Adiyaman: Nemrut Da., 2140 m, Sorger 73-29-6! C9 Hakkari: 28 km S.E. of Başkale, 2000 m, 20 v 1966, Eiselt! C10 Hakkari: Nehil Çayi, 62 km from Hakkari to Yüksekova, 1750 m, D. 44919! Soviet Armenia, N. Iraq, N.W. & W. Iran. lr.-Tur. element. A very variable species, previously divided into numerous varieties, subvarieties and forms. In my opinion, such division seems unjustified. Intermediates link many of the taxa, which frequently grow close together, and correct determination of infraspecific taxa is consequently often difficult. Forms previously classified as R. lanata var. punctata, and R. lanata var. canescens subvar. detonsa are ± glabrescent with only calcified tubercles on the bright green leaves; they are very distinct and may warrant varietal recognition. On the other hand it is probable that they are only calcicole modifications. Collateral cultivation would be desirable in order to clarify their taxonomic status. Vars. lanata and canescens are maintained here as they are quite readily identified by the shape of the basal and lowest cauline leaves. Although their ranges overlap considerably, intermediates appear to be rare. |