| Brassicaceae |
|---|
 DAVIS var_ subradiata DESV_jpg.jpg)
Isatis cappadocica DESV. subsp. subradiata (RUPR.) DAVIS var. subradiata DESV. |
 Isatis cappadocica DESV. subsp. subradiata (RUPR.) DAVIS var. subradiata DESV. |
Isatis L. |
| I. cappadocica Desv. in J. Bot. Appl. 3: 174 (1814). Syn: /. latisiliqua sensu Boiss., Fl. Or., 1: 377 (1867) p.p. non Stev. Perennial, often suffrutescent. Stems 20-70 cm. Leaves thin, like the stems usually pubescent or glabrous; basal leaves oblanceolate or obovate, entire or weakly dentate; median stem leaves oblong to linear-oblong or oblong-lanceolate, obtusely or acutely auricled. Flowers 4-5 mm. Fruits 10-30 x 5-20 mm, in Turkey obovate, elliptic, ovate, suborbicular, ovate-rhomboid or cuneate-oblong, shortly and densely hairy or glabrous, wing thin, usually much broader than the loculus, flat or undulate, loculus not transversely wrinkled, midrib slender. An extremely polymorphic species whose limits and subdivision remain obscure. As circumscribed here, the species is equivalent to most of Boissier's inter-pretation of /. latisiliqua Stev. though it excludes /. latisiliqua itself, a distinctive species confined to Caucasia and distinguished from /. cappadocica by its hispid stem bases and transversely wrinkled loculus (as in Sameraria armena (L.) Desv.). The key below attempts to cope with most of the major variants found in Turkey; other taxa in the same complex occur in Caucasia (/. nummularia Trautv., /. karjaginii Schischk.) and Iran (/. besseri Trautv., /. stenophylla Bornm. & Gauba, etc.). Examination of the type in Paris shows /. cappadocica to be the earliest name for the group. 1. Fruits 10-12x5-9 mm, usually ovate-rhomboid, flat, densely velutinous or glabrous; stems 20-30 cm; basal leaves obovate or obovate-oblanceolate subsp. cappadocica 1. Fruits 12-30x8-20 mm, elliptic, ovate, obovate or suborbicular, flat or undulate, puberulent or glabrous; stems 30-70 cm; basal leaves oblanceolate, often narrowly so 2. Wing of fruit strongly undulate when ripe 3. Fruit broadly elliptic to suborbicular, 13-17 x 9-17 mm subsp. steveniana 3. Fruit ovate-elliptic, 20-30 x 12-20 mm, often mottled with purplish brown subsp. macrocarpa 2. Wing of ripe fruit nearly flat 4. Fruits cuneate-oblong, 24-26 x8-9 rnm, tapered below, wing as broad as loculus subsp. nurihakensis 4. Fruits suborbicular, obovate, ovate or elliptic, 12-30 x 9-20 mm, wing manifestly broader than loculus 5. Fruit suborbicular, 12-16 x 12-İ6 mm (sometimes rounded-elliptic), obtuse or emarginate (inflorescence of short arcuate branches; plant suffruticose) subsp. alyssifolia 5. Fruits obovate, elliptic or ovate-elliptic, 14-30 x9-20 mm 6. Fruits usually obovate or elliptic, 14-24 x 9-12(-17) mm, acute at the base or at both ends; stem leaves usually obtusely auricled 7. Fruits obovate, sometimes broadly elliptic (rarely ovate-elliptic), usually obtuse and less than 2 x longer than broad, puberulent or glabrous subsp. subradiata var. subradiata 7. Fruits elliptic, usually 2 x longer than broad, obtuse or emarginate, tomentose-puberulent or glabrous subsp. subradiata var. gudrunensis 6. Fruits ovate-elliptic, 20-30 x 12-20 mm, rounded at least at the base, often mottled with purplish brown; stem leaves subacutely auricled subsp. macrocarpa subsp. subradiata (Rupr.) Davis in Notes R.B.G. Edinb., 26: 18 (1964). var. subradiata. Figure 15, p. 291. Syn: /. subradiata Rupr., Fl. Cauc, 131 (1869); /. latisiliqua Stev. subsp. subradiata (Rupr.) Bordz. in Izv. Kiev Bot. Sada, 12-13: 112 (1931). Fl. 5? Banks, oak scrub, etc., 1450-1750 m. Type: Caucasus, ad fl. Wjera, pr. Tiflis (LE!). Very scattered. B9 Van: 5 km N of Çatak, 1750 m, D. 23268 (robust form with stem hirsute below)! C4 Konya: Taşkent, 1450 m, Hub.-Mor. 9236 (fruit exceptionally broad)! C5 Seyhan: d. Feke, Görümse, 1600 m, Kotschy suppl. 170. Transcaucasia, N. Iraq, W. Iran. Ir.-Tur. element. |