O. spinosa L., Sp. PL 716 (1753).
Perennial with erect or ascending stems, without stolons, 30-60 cm, spinescent, with an indumentum of short pedicellate or sessile glands and longer eglandular hairs, sometimes glabrescent. Upper leaves often unifoliolate, the others tri-foliolate; leaflets ovate to oblong, denticulate, 5-20 x 3-10 mm. Flowers shortly pedicellate, mainly single in the leaf-axils, forming loose racemes. Calyx cam-panulate, 5-10 mm, parted to ?-¾ into linear-lanceolate 1-3-nerved lobes; teeth glabrescent or covered with sessile to subsessile glands, sometimes intermixed with longer eglandular hairs. Corolla pink, 6-10(-15) mm, occasionally white, standard often with purple stripes. Legume ovate, glandular-hairy, 1-3-ovuled. Seeds orbicular, tubercled or smooth, 2 mm.
1. Seeds tubercled subsp. antiquorum
1. Seeds smooth subsp. leiosperma
subsp. leiosperma (Boiss.) Širj. in Beih. Bot. Centr. 49(2):590 (1932). Syn: O. leio-sperma Boiss., Fl. Or. 2:57 (1872). Fl. 5-8. Stony slopes, vineyards, cultivated land, s.l-2250 m.
Type: [Turkey C6 Hatay] Dispersa locis argillosis montis Amani prope Beilan (Belen), 600 m, v, vi 1862. Kotschy 296 (holo. G!).
Widespread. A1(A) Çanakkale: Assos, Sint. 1883:951! A2(E) Istanbul: Kücük Çekmece, B. Post! A3 Kocaeli: Sapanca G., 50 m, D. 36213! A4 Kastamonu: above Küre, 1200 m. D. 38746! A5 Amasya: Amasya, Maniss. 83b! A7 Gümüşane: Janur De., Sint. 1889:6116. A8 Erzurum: Olti-çay, Radde. Bl Izmir: Izmir, Unger. B4 Ankara: Beynam 55 km S W of Ankara, Ledingham & Ekim 4329! B6 Maraş: Distr, Göksun, Hobek Da., 1700 m, D. 20184! B7 Elaziğ: Maden. S W of Devedüzü, Romieux! B8 Muş: Varto, 1560 m, D. 46298! B9 Van: Warak Da. nr. Van, 2200 m, Náb. 3016. C2 Denizli: Pamukkale, Reese! C3 Antalya: nr. Finike, 1350 m, Khan et al. 251! C5 Adana: Feke, 600 m, D. 19647! C6 Gaziantep: Dülük Baba, 900-1200 m, Haradj. 1352! C7 Urfa: Urfa to Akçakale, 450 m, D. 28114! C8 Mardin: Khurs, Sint. 1888:1038! C9 Hakkari: Koçanis, 2250 m, D. 24324! Is: Kos, Rech. 7962! Kalimnos, Rech. 7849!
Balkans, Cyprus, W. Syria, N. Iraq, N. & W. Iran. Intermediate forms between O. arvensis and O. spinosa subsp. leiosperma are frequent around Istanbul. They have been described as O. decipiens Azn. (Mag. Bot. Lap. 2:138, 1903), as O. spinosa ssp. antiquorum var. glandulifera Širj. (Gen. Ononis rev. crit. 586, 1932) and as O. repens var. pichleri Širj. (op. cit. 601). The following gatherings may be cited: A2(E) Istanbul: Yedikule to Makriköy, Azn.! Kilyos, B. Post ! A2(A) Bursa: Bursa, 1874, Pichler (type of O. repens var. pichleri)! A3 Zonguldak: Kozlu, Khan et al. 814! A6 Samsun: Samsun, Tobey 5/9! Other gatherings col-lected near Istanbul may be considered as intermediate between O. repens L. and O. spinosa subsp. leiosperma: A2(E) Istanbul: Ayimana to Makriköy (Bakirköy), Azn. 586 bis! A2(A) Istanbul: Küçük su valley, B. Post! Bostanci, Azn.! ('O. byzantina' Azn. Hb.). |