| Fabaceae |
|---|
 P_ W_ BALL _jpg.jpg)
Vicia villosa ROTH subsp. eriocarpa (HAUSSKN.) P. W. BALL |
Vicia L.
|
| V. villosa Roth, Tent. Fl. Germ. 2:182 (1795). Annual or biennial, villous or adpressed-pilose to almost glabrous, 15-9O(-12O) cm, climbing or procumbent. Leaflets 4-10-paired, 0.3-3.5 cm, oblong-ovate to narrowly linear; stipules semi-hastate or sagittate to lanceolate; tendrils branched. Peduncle as long as the leaf or shorter, 3-40-flowered. Pedicels l-2(-3) mm. Flowers 12-22 mm, violet-blue or ± blue. Calyx 5-8 mm, strongly gibbous, with an oblique mouth, villous to sparsely pilose-pubescent; teeth shorter to longer than tube, narrowly subulate to lanceolate-triangular, ciliate or glabrescent. Limb of standard | to nearly as long as the claw; keel navicular. Style laterally subcompressed. Legumes oblong or oblong-rhomboid. (l.5-)2-3(-3.5) × (0.5-)0.6-1(1.2) cm, glabrous or pubescent. Seeds (l-)2-8, hilum 1/6-1/12 of perimeter. A very polymorphic species. Four overlapping subspecies (as recognised in Fl. Eur. 2:132) occur in our area; the taxonomic situation is particularly puzzling in W. Anatolia and the neighbouring islands. 1. Stems ± villous; leaflets usually 7-10-paired, elongate, never obtuse; stipules up to 5 mm broad: raceme 10-30-flowered; lower calyx tooth at least as long as tube, ciliate; limb of standard at least f as long as claw; legume glabrous subsp. villosa 1. Stems glabrescent or adpressed-pilose; leaflets 4-10-paired; stipules less than 3 mm broad: racemes (3-)5-15-flowered; lower calyx tooth generally shorter than tube, usually subglabrous; limb of standard 1/2 as long or nearly as long as claw: legume glabrous or pubescent 2. Leaflets linear-oblong, (4-)6-10-paired, some often longer than 17 mm; raceme 5-15-flowered; standard violet, limb 1/2-2/3 as long as claw 3. Ovary and mature legume glabrous subsp. dasycarpa 3. Ovary and legume persistently pubescent subsp. eriocarpa 2. Leaflets ± elliptic, 4-7-paired, shorter than 17 mm; raceme 3-7-flowered, lax; corolla 14-22 mm, ± blue, often yellow-tinged, standard at least § as long as claw (legume glabrous or pubescent) subsp. microphylla subsp. eriocarpa (Hausskn.) P. W. Bail in Feddes Rep. 79:45 (1968). Figure 3, p. 323. Syn: V. varia Host var. eriocarpa Hausskn. in Mitt. Thür. Bot. Ver. n.s. 5:87 (1893); V. salaminia Heldr. & Sart. var. stenophylla Hausskn. in Mitt. Thür. Bot. Ver. n.s. 5:88 (1893); ?V. salaminia Heldr. & Sart. subsp. macro-phyllaha Cand. in Bull. Soc. Bot. Fr. 45:187 (1898); V. eriocarpa (Hausskn.) Hal., Consp. Fl. Graec. 1:489 (1900). Fl. 3-7. Pinus forest, macchie, hedges, banks, fields, nr. water, s. 1.-1750 m. Syntypes: [Greece] in dumosis ad radicem Hymetti: in collibus ad Akro- et Neo-Korinthum, supra Naupliam; in P.D. reg. infer. pr. mon. Korona, [Hauss-knecht]. Mainly Turkey-in-Europe, N.W., W. & S. Anatolia, Islands. A1(E) Edirne: 5 km S E of Bahçeköy (3 km ESE of Çamlica), Kuru Da.. Bauer & Spitz. 557! A1(A) Çanakkale: Dardanelles, in Rhodus valley, Sint. 1883:46! A2(E) Istanbul: Kiathane to Şişli, 16 vi 1893, Azn.! A2(A) Istanbul: Kartal, iv 1913, B. Post! A3 Bolu: Düzce to Akçakoca, 250 m, D. 37476! B1 Izmir: Kaeşiya (?), 50 m, A. Tanoy 2446! B2 Uşak: 6 miles from Uşak to Sivasli, 900 m, Coode & Jones 2419! B4 Ankara:Orman Çiftlik, Üresin 182 (introduced?)! Cl Izmir: d. Kuşa-dasi, Samsun Da., Kayacık & Yalt. 3440! C2 Muğla: Marmaris, D. 41428! C3 Antalya: d. Elmali, Üçkuyular, 1750 m, Khan et al. 268! C4 Içel: d. Anamur, foot of Kalandöken Da., 100 m, D. 25935! Is: Lesvos, Rech. 5749! 5766! 5500! Khios, Platt 444! Samos, Rech. 3588! Kos, D. 40504! Rodhos, Rech. 7034! 7206 (approaching subsp. microphylla)! Greece, Aegean, W. Syria, Cyprus, Iran, N. Iraq. It seems probable that V. salaminia Heldr. & Sart. subsp. macrophyllaria Cand., described from Lesvos, provides the earliest epithet for this plant at subspecific rank, but as Candargy's type has been destroyed we have preferred to adopt the name subsp. eriocarpa. This subspecies largely replaces subsp. dasycarpa in much of S.W. Asia (cf. note under that subspecies). Intermediates occur. |