Lamiaceae

N. nuda L., Sp. P1. 570 (1753).
Perennial; stems several, erect, 20-90 cm, finely adpressed eglandular-pilose, villous, with glandular hairs or papillae or rarely subglabrous. Leaves ovate to ovate-oblong, 1.5-5(-7) x l-2.5(-3.5) cm, adpressed pubescent with numerous sessile glands, crenate, cordate or truncate; petiole 0.5-2 cm. Inflorescence paniculate of numerous many-flowered verticillasters, rarely thyrsoid. Bracteoles shorter than calyx tube or subequal to it, linear to linear-lanceolate. Calyx tubular in flower, 5-7 mm, ± ovoid in fruit, yellowish-green or violet-blue, tube and mouth straight, adpressed eglandular-pilose with sessile glands, with glandular hairs or papillae, or with spreading eglandular hairs; teeth erect, 2-3 mm, blunt, separated by rounded sinuses, often with membranous margins. Corolla white, cream or violet-blue, 7-10 mm; tube slightly curved. Nutlets ovoid oblong, trigonous, c. 2 x 1 mm, tuberculate, prominently so at apex with crowded projecting tubercles.
One of the commonest and most widespread species in Turkey. It varies considerably in flower colour, indumentum, inflorescence and calyx characters. Much of the variation is at the local population level, but with the considerable amount of material available it seems that four subspecies are worth recognition.
1. Inflorescence axis, calyces and corollas ± violet-blue subsp. nuda
1. Inflorescence axis, calyces and corollas green, yellowish-green or white
2. Indumentum on stem with numerous short capitate glandular hairs subsp. glandulifera
  2. Indumentum on stem without or with very few, capitate glandular hairs
   3. Bracts as long as calyx tube; indumentum on stem of spreading tangled hairs subsp. lydiae
   3. Bracts shorter than calyx tube; indumentum of very short pilose ± adpressed hairs (occasionally densely hairy) subsp. albiflora
subsp. albiflora (Boiss.) Gams in Hegi,I11. Fl. Mittel-Eur. 5(4): 2372 (1927). Syn: N. sulphurea C. Koch in Linnaea 21:677 (1848); N. marrubioides Boiss. & Heldr. in Boiss., Diagn. ser. 1(12):67 (1853)! N. nuda L. var. albiflora Boiss., Fl. Or. 4:663 (1879)! N. lycia Stapf in Denkschr. Akad. Wiss. Wien, Math.-Nat. Kl. 50(l):99 (1885)! TV. nawaschini Bordz. in [Mem. Soc. Nat. Kiev] Protok. Zased. Kievsk. Obshch. Estest. 25(1): 109 (1907); N. tolypantha Stapf var. bithynica Bornm. in Beih. Bot. Centr. 24(2):489 (1909)! TV. nuda L. subsp. marrubioides Hub.-Mor. & Davis in Kew Bull. 1951:83 (1951)! Map 40. Fl. 5-8. In deciduous and coniferous woodland, meadows, stream-sides, rocky limestone and grassy slopes, fieldsides, 850-2750 m.
Numerous syntypes from Greece, Turkey, Caucasus, Persia and Lebanon.
Throughout most of Anatolia but absent from parts of the N.W. & W. A2(A) Bursa: Ulu Da., Aras, A. & T. Baytop (ISTE 2459)! A3 Bolu: above Mudurnu, 1250 m, D. 37090! A4 Çankiri: Ilgaz Da., 1610 m, Hub.-Mor. 15585! A5 Kastamonu: Tosya to Yaçillar köyü, Karanlikdere, Kilinç 3685! A6 Amasya: Erbaa above Kozlu, 850 m, Tobey 2200! A7 Giresun: 22 km N. of Şebinkarahisar, 1700 m, Sorger 69-29-21! A8 Rize: d. Hemşin, Ortaköy, 2300 m, D. 21248! A9 Kars: Sarikamiş, 2100 m, D. 30803! Bl Balikesir: mt. Ida (Kaz Da.), Sint. 1883:646! B2 Kütahya: Demirci to Simav, Simav Da., 1340 m, Hub.-Mor. 12768! B3 Konya: Akşehir, 1000 m, Bornm. 1899:5480! B4 Ankara: Elma Da. nr Ankara, c. 1700 m, 25 vi 1932, Kotte! B5 Adana: Bakirdağ to Saimbeyli, 1640 m, Hub.-Mor. 11327! B6 Maraş: d. Göksun, Bin-boğa Da., Isik Da., 1900 m, D. 20072! B7 Tunceli: Pertek to Tunceli, 1100 m, D. 29151! B8 Muş: Gümgüm (Varto), Merga Sauk, 1830 m, Kotschy 1859: 368! B9 Van: Başkale to Hoşap, 2350 m, Hub.-Mor. 11328! C2 Antalya: 31 km S.W. of Elmali, 1150 m, Sorger 65-25-27! C3 Isparta: Dedegöl Da., 1600 m, Sorger 65-43-54! C4 Içel: E. of Gülnar, c. 950 m, Hub.-Mor. 10165! C5 Adana: d. Karaisalı, Osmancik Y., 1450 m, Hub.-Mor. 15586! C6 Hatay: Hasanbeyli, 1370 m, Haradj. 2240! C7 Adiyaman/Malatya: Ak Da., Malatya to Kahta, 2200 m, Hand.-Mazz. 2352! C9 Hakkari: Kara Da., 2750 m, D. 24325! C10 Hakkari: 26 km from Şemdinli to Yuksekova, 2000 m, D. 45046!
S. Jugoslavia, N. Greece, Transcaucasia, N., N.W. & C. Iran, N. Iraq, Lebanon. European specimens generally have fewer flowers in the verticil-lasters and are generally less sturdy in appearance than the Turkish plant; they also appear to be more frequently male sterile.