Lamiaceae

S. frigida Boiss., Diagn. ser. 1(5): 10 (1844). Syn: S. spinulosa Montbret & Aucher ex Bentham in DC, Prodr. 12:283 (1848)! S. frigida Boiss. var. oblongifolia Boiss., Fl. Or. 4:621 (1879)! S. frigida Boiss. var. albiflora Bornm. in Beih. Bot. Centr. 24(2):487 (1909)! Map 54.
Perennial herb with a thick woody rootstock. Stems solitary or several, erect, 10-30(-50) cm, below pilose to villous with sessile glands, densely glandular above with capitate glands. Leaves mostly basal, variable, ovate to narrowly oblong, 2-12 x 1.2-5 cm, crenulate to erose, rugulose, arachnoid to ± lanate with many sessile glands; petiole 1.5-8 cm. Inflorescence little branched to widely paniculate. Verticillasters 2-6-flowered, usually distant. Bracts ovate to orbicular, c. 8-12 x 7-11 mm. Pedicels 2-3 mm. Calyx campanulate to infundibular, 7-10 mm, to 10-12 mm in fruit and widening, densely capitate-glandular; teeth usually prominently spinulose; upper lip tridentate, median tooth much shorter, truncate or not. Corolla white to lilac, 12-16 mm; tube c. 8 mm, ventricose, squamulate; upper lip narrow, somewhat falcate, scarcely bifid. Stamens B. Nutlets ovoid, c. 3.5 x 2.5 mm. 2n = 20, 21, 22. Fl. 5-7. Pinus nigra, Juniperus, Abies and Pyrus woodland, limestone slopes and crevices, meadows, 1000-2450 m.
Type: [Turkey C2 Denizli] in regione alpina excelsiori montium Cariae, Cadmi orientalis (Honaz Da.) supra Colossam (Honaz) et occidentalis (Baba Da.) supra Gheyra (Göyre), vi 1842, Boissier (holo. G!).
Inner & S.W. Anatolia. A4 Ankara: Çamkoru, N.W. of Kizilcahamam, 1350 m, Bozakman & Fitz 807! B2 Kütahya: Murat Da. above Banaz, 2000 m, Coode & Jones 2499! B3 Eskişehir: Sündiken Da., c. 1000 m, Ekim 632! B4 Konya: 90 km N. of Konya, M. Zohary & Orshan 281211 (doubtful deter-mination)! B5 Kayseri: Bakir Da. above Kisge, 1900 m, D. 19443! B6 Sivas: Gürün to Sivas, 1710 m, Hub.-Mor. 11865! B7 Malatya: 5 km E. of Malatya, 1300 m, Hub.-Mor. 8988! B8 Erzincan: 17 km from Tercan to Aşkale, 1680 m, Hub.-Mor. 10768! B9 Bitlis: Bitlis to Tatvan, 1900 m, D. 22354! C2 Denizli: Honaz Da., 1300 m, Sorger 62-32a-29! C3 Isparta: Davras Da., A. Baytop (ISTE 9620)! C4 Konya: Bozkir to Çamli, 1400 m, Çetik et al. 1017! C5 Içel: Tekir Tepesi, 1270 m, Sorger 62-68-37!
N.W. Iran. Ir.-Tur. element. Close to S. modesta and S. yosgadensis. With some affinities to S. argentea and S. microstegia but with much smaller cor-ollas. A most variable species in all respects. Although there is a core of speci-mens which is fairly typical for the species, there are many individuals which because of characters of habit, indumentum, leaf shape, bract size and calyx shape are anomalous. Mostly they occur in the northern peripheral parts of the range (e.g. A4, B2, B3). Male sterile sex-forms are very common and these plants tend to be aberrant in other respects. No definite hybrids have been reported but D. 19915 (B6 Maraş; d. Göksun, Yalak, 1400 m) combines fea-tures of S. frigida (calyx and corolla) and S. microstegia (height and indu-mentum) and is not readily assignable to either.