Lamiaceae

T. chamaedrys L., Sp. Pl. 565 (1753). Rech. fil. in Bot. Archiv 42:338-389 (1941). Syn: T. officinale Lam., Fl. Fr. 2:414 (1779); T. pseudochamae-drys Wender. in Flora 9:358 (1826). Ic: Bot. Archiv 42:417-420 (1941).
Suffrutescent perennial herb, often rhizomatous, 5-50 cm, with very variable indumentum. Leaves oblong or obovate-oblong, cuneate at base, usually crenate-dentate or shortly lobulate, lobes entire or toothed. Verticillasters (2-)4-8-flowered, in dense or ± lax terminal racemes, floral leaves (bracts) usually distinct from cauline leaves, entire or toothed; pedicels equalling or usually shorter than calyx. Calyx tubular-campanulate, subgibbous at base, teeth lanceolate-triangular, ± subulate-tipped, margins usually densely short-haired and sparsely long-hairy, or only long-hairy, or glabrous, usually purplish. Corolla reddish-purple, 2 x calyx. Stamens at least as long as upper lateral lobes
1. Bracts all overtopping calyces
  2. Leaves subglabrous or shortly hispid beneath subsp. lydium
  2. Plant bearing long eglandular or glandular hairs
   3. Calyx teeth glabrous or sparsely long-haired subsp. trapezunticum
   3. Calyx teeth densely covered with short and long hairs subsp. chamaedrys
1. Bracts mostly shorter than calyces
    4. Plant shortly retrorse- and crispate-hairy; stem hairs diminishing in length downwards subsp. sinuatum
    4. Indumentum not as above
     5. Upper bracts entire, all ciliate-margined; stem hairs not diminishing in length upwards;leaves ± discolorous subsp. syspirense
     5. Bracts all dentate, not usually ciliate-margined; stem hairs diminishing in length upwards;leaves concolorous subsp. tauricolum
subsp. syspirense (C. Koch) Rech. fil. in Ann. Naturh. Mus. Wien 51:427 (1941). Syn: T. syspirense C. Koch in Linnaea 21:704 (1848); T. chamae-drys L. var. canum Boiss., Fl. Or. 4:816 (1879) p.p. non T. canum Fisch. & Mey. (1835); T. chamaedrys L. var. hirsuta Čelak. in Bot. Centr. 14:220 (1883); T. chamaedrys L. var. gymnobotrys Celak. op. cit. 221 (1883)! Fl. 6-8. Limestone or igneous rocky slopes, steppe, 90-2430 m.
Type: [Turkey A8 Erzurum] Horasan, im Gaue Sber (Ispir), sehr haufig auf porphyr, 915-2430 m, [1844], C. Koch.
Inner & adjacent N. & S. Anatolia. A4 Kastamonu: 5 km W. of Kastamonu, 900 m, D. 38771! A5 Sinop: Gerze, 30 km S. of Sinop, Hub.-Mor. 16358! A6 Amasya: Erbaa to Koçak, 300 m, Tobey 2177! A7 Gümüşane: Gümüşane, Sint. 1894:6099! A8 Erzurum: 20 km N. of Aşkale, 1900 m, Kukkonen 8053! B2 Kütahya: 18 km S. of Kütahya, 1000 m, D. 36935! B3 Afyon: Sultan Da., Bornm. 1899:5422. B4 Ankara: Beynam forest, D. 13013! B5 Kayseri: Talas, 1230-1270 m, Hub.-Mor. 10794! B6 Maraş: 5 km W. of Elbistan, 1070 m, Hub.-Mor. 12579! B7 Malatya: 3 km E. of Malatya, 950 m, Hub.-Mor. 8956! Sivas: Divrik (Divriği), c. 1000 m, Bornm. 1893: 3516 (type of T. ixodes, photo!). B8 Erzurum: Palandöken Da., 2300-2350 m, D. 47352! B9 Van: Toprakkale, 1620-1800 m, Hub.-Mor. 8958! C2 Antalya: d. Elmali, Bey Da., Çamçukuru Y., 1750 m, Khan et al. 324! C3 Burdur: 5 km W. of Burdur, 900 m, Hub.-Mor. 8402! C4 Konya: Sara nr Ermenek, 1872, Péronin 186 (type of var. gymnobotrys)! C7 Malatya: 74 km S.W. of Malatya, 1550 m, Sorger 71-45-27!
Transcaucasia, N. Iran & Khorassan. Ir.-Tur. element. This rather distinctive subspecies differs from the others in its discolorous leaves, and usually entire bracts which (except the lowest pair) are equal to or shorter than the flowers; T. ixodes Bornm. is only a glandular variant. It is possible to see many intermediate forms between this subspecies and subspp. tauricolum, chamaedrys and trapezunticum, mainly in contact areas; they are listed at the end of the species.