Rosaceae

R. heckeliana Tratt., Ros. Monogr. 2:85 (1823).
Low-growing shrub 0.l-0.6(-l.5) m, many-branched, ± twisted with creeping underground stems. Cortex of young shoots ± pubescent. Prickles straight or slightly curved, up to 1.1 cm long, thin and bristle-like, ± uniform, often ascending, often pubescent at least on young branches, sparse. Leaflets 5-7, elliptic to suborbicular, 1-7 x0.8-3.7 cm, acute to obtuse or retuse, with a rounded to cun-eate base, shortly adpressed-pubescent or velutinous, usually eglandular, pale greyish-green, usually uniserrate, teeth distinct; rachis densely pubescent, ± prickly and with stalked glands; stipules broad, pubescent, entire or glandular-serrulate, auricles diverging. Flowers solitary or 2-3 together. Pedicels 0.5-1.5(-3) cm, densely pubescent and often with some stalked glands. Outer sepals usually lobed, narrowly ovate with a long narrow apex dilated above, 1-2.5 cm, eglandular or with sparse glands on the back, ascending to erect after anthesis, persistent until the fruit matures. Petals 1.1-1.5 x1-1.2 cm, usually pale to deep pink, rarely white, often persistent after anthesis. Styles lanate, stigma-head compact, semiglobose, almost as broad as the narrow, ± concave disc, orifice wide. Hypan-thia subglobose to ovoid, up to 1.5 cm diam., smooth or usually densely setose, setae often long, not very stiff, glandular or eglandular, dull to scarlet red. Fl. 6-7. In scrub or open places on rocks, slopes and screes or at wood edges, usually on limestone, 1300-2900 m.
1. Prickles rather stout with a broad compressed dilated base, glabrous; leaflets elliptic to ovate, usually acute and with a cuneate base, green and ± pubescent above, pale greyish-green and velutinous beneath; stipule auricles rather long, acute; sepal lobes elliptic, apex much dilated; hypanthia ovoid subsp. vanheurckiana
1. Prickles often weak, acicular, usually with a small rounded base, ± pubescent, sparse; leaflets ovate to suborbicular, usually obtuse to retuse with a rounded base, pale greyish-green and velutinous on both surfaces; stipule auricles shortly triangular; sepal lobes small, linear-lanceolate, sepal apex only slightly dilated; hypanthia ± subglobose subsp. orientalis
subsp. vanheurckiana (Crépin) Ö. Nilsson, comb. et stat. nov. Syn: R. vanheurck-iana Crépin in Boiss.. Fl. Or. 2:683 (1872)! R. coriifolia Fries var. vanheurckiana (Crépin) Christ in Boiss., Fl. Or. Suppl.: 216 (1888); R. atropatana Sosn. in Izv. Akad. Nauk Azerb. S S R 10:22 (1944); R: hakkariensis Manden. in Notes R.B.G. Edinb. 30:337(1970)!
Syntypes: [Turkey B8] in alpibus prov. Musch (Muş) Armeniae australis ad 1830 m, Kotschy 789; in valle Merga Sauk, 2130 m, Kotschy 369 (as R. orientalis). E. Anatolia. B8 Bingöl: nr. Boglan, 1370 m, Kotschy 540! B9 Ağri: d. Suluçem (Musun), S. end of Balik G., 2300 m, D. 47264! Bitlis: Süphan Da., above Adilcevaz, 2900 m, D. 24704! Van: Çuk Gediği pass between Hoşap and Başkale, 2600 m, Birand & Karamanoğlu 486! C9 Hakkari: Cilo Da., between Diz deresi and Cilo Y., 2440 m, D. 24247 (type of R. hakkariensis Manden.)!
Soviet Armenia, W. Iran. Ir.-Tur. element. The type of 'R. hakkariensis' is atypical in being less pubescent and in having more rounded leaflets with some glands, longer pedicels and ascending sepals; it may be a hybrid.