Rosaceae

R. pulverulenta Bieb., Fl. Taur.-Cauc. 1:399 (1808). Syn: R. glutinosa Sm. in Sibth. & Sm., Prodr. Fl. Graec. 1:348 (1809); R. poteriifolia Schott & Kotschy in sched.! R. libanotica Boiss., Diagn. ser. l(10):4 (1849) pro parte! Ic: Sibth. & Sm., Fl. Graeca 5: t. 482 (1825); Fiori, Ic. Fl. Ital. f. 1811 (1899).
Low-growing shrub, 0.l-0.5(-l) m, often forming dense patches or mats. Prickles weak to rather coarse, straight to curved, occasionally hooked, with a ± dilated, usually acentric base, usually passing into and dispersed with a dense covering of acicles, bristles and stalked glands. Leaves often small, ± sticky, aromatic; leaflets usually 5, ovate, elliptic or suborbicular, refuse or subacute, with a rounded to cuneate base, glabrous to ± pubescent, usually very glandular, at least beneath, glandular-biserrate, teeth short, 8-20 on each side; rachis ± pubescent, glandular and often prickly; stipules glandular, rather broad with diverging, ± triangular, short auricles. Flowers solitary or 2-4 together. Pedicels 0.2-1.5(-2.5) cm, glabrous or pubescent, usually ± densely stalked-glandular, only rarely smooth. Sepals ovate with a rather short, often not much dilated apex, up to 2.5 cm long, outer sepals pinnatifid with lanceolate, glandular-biserrate lobes, ± glandular on back, patent to erect after anthesis, usually not deciduous before the fruit matures. Petals 1-1.8 cm long, pink. Styles villous to lanate, stigma-head ± compact, depressed-semiglobose;disc often very narrow, orifice wide. Hypanthia globose to ovoid, 1-2 cm long, smooth or ± glandular-hispid, scarlet. Fl. 6-7. Dry open, often grazed places, Juniperus scrub, Picea woodland, Artemisia steppe, stony and rocky slopes or banks, screes, subalpine meadows on limestone or igneous substrate, 700-2550 m.
Type: [Caucasus] in collibus circa acidulam Narzana Caucasi subalpini, Bieberstein (LE, photo E!).
Throughout Anatolia. A2(A) Bursa: Bithynian Olympus (Ulu Da.), 1874, Pichler! A3 Bolu: Ala Da., on Kartal Kaya Tepe, 2100-2200 m, D. 37338! A4 Ankara: 26 km N. of Kizilcahamam, 1700 m, Sorger 70-5-81! A5 Amasya: Sana Da., above Yasian, Bornm, 1890:2858! A7 Gümüşane: Gümüşane, 1220 m, Balls 1985! A9 Kars: N. side of Akçay to Cumaçay pass, E. of Kağizman, 2350 m, D. 46762! W. of Kars, 1700 m, D. 32582! B1 Balikesir: Mt. Ida (Kaz Da.), nr. Kareikos, Sint. 1883:480! Kapu Da., Sint. 1883:480c! B2 Izmir: Boz Da., 1600 m, Sorger 68-16-113! B3 Konya: Sultan Da., 1750 m, Bornm. 1899:4432! B5 Kayseri: Erciyas Da., E. slope, 1800 m, M. Zohary & Plitm. 17600-8! B6 Sivas: 42 km N. of Gürün, 1700 m, Sorger 70-33-62! B7 Malatya: 56 km from Gölbaşı to Malatya, Alava 7028! B8 Bitlis: Kambos Da., above Kotum, 1980 m, D. 22218! B9 Bitlis: Kotum, Karz Da. above Kamer, 2300 m, D. 24601! B10 Kars/Ağri: Mt. Ararat (Ağri Da.), Szovits! C2 Muğla: Girdev Da., above Duger, 1600 m, D. 13810! C3 Isparta: Davras Da., 1500 m, A. & T. Baytop 4277! C4 Konya: Karanje De., Geyik Da. to Bozkir, 1850 m, D. 14618! C5 Niğde: Ala Da., Narpiz gorge, 2450 m, Wood & Gibson 242! C5/6 Taurus Mtns., Kotschy 186 (type of R. poteriifolia)! C6 Maraş: Andirin, Mazgaç Tepesi, 2200 m, 1958, Markgraf! C7 Adıyaman: Nemrut Da., Hand.-Mazz. 2090. C9 Hakkari: Koçanis, 2300 m, D. 24312! Is: Khios, Platt 219! Samos, mt. Kerki, nr. Hag. Ilias, Fors.-Maj. 379!
S.E. Europe, W. Syria, Cyprus, Caucasus, Soviet Azerbaijan, Soviet Armenia, Iran, Afghanistan. A variable species with regard to growth, armature and indumentum. Of the specimens cited above, that from B1 Balikesir {Sint. 1883:480c) differs in its small sparse prickles and absence of glands on the stem cortex and fruit; that from A9 Kars (D. 32582) in its long pedicels, and that from B1 Balikesir {Sint. 1883:480) in its hooked prickles. Mandenova (op. cit. p. 339) treated tall growing forms with large leaves from Easternmost Anatolia as R. pulverulenta sensu stricto and the remainder as R. glutinosa; transitional forms occur however in different parts of Turkey so that it is impossible to maintain the two as distinct species. The following specimens are of hybrid origin; R. pulverulenta x R. canina-C9 Hakkari: 6 km from Hakkari to Kaval (Piyanis), 2050 m, D. 45428! R. pulverulenta x R. dumalis subsp. boissieri-B7 Erzincan: Sipikör, Kainik Dere, Sint. 1890:3153!