Scrophulariaceae

R. angustifolius C.C. Gmelin, Fl. Bad. 2:669 (1806).
subsp. grandiflorus (Wallr.) D.A.Webb in Bot. J. Linn. Soc. 65:269 (1972). Syn: Alectorolophus grandiflorus Wallr., Sched. Crit. Fl. Hal. 316 (1822); A. ponti-cus Sterneck in Abh. Zool.-Bot. Ges. Wien 1(2):48 (1901); A. vernalis Zinger in Trudy Tbilissk. Bot. Sada 12: 182 (1913); Rhinanthus vernalis (Zinger) Schischkin &Sergievsk.in Krylov, Fl. Zap. Sibir. ed. 2, 10:2530<1939); R. colchicus Vassilcz. in Not. Syst. (Leningrad) 17:378 (1955); R. ponticus (Sterneck) Vassilcz. in Fl. URSS 22:667 (1955); R. angustifolius C.C.Gmelin 'grex' grandiflorus (Wallr.) Soóin Acta Bot. Acad. Sci. Hung. 16:203(1970). Ic: Hegi, III. Fl. Mittel.-Eur. ed.
2, 6(1):394, f. 193 (1972). Figure 24, p. 771.
Stems to 70 cm, erect, usually branched above, eglandular or glandular pubescent or villous. Leaves narrowly oblong-lanceolate, 2.5x0.5-6x2 cm, crenate-dentate, with revolute margins. Flowers borne in upper leaf axils, forming compact or loose spikes. Bracts leaf-like, ovate, longer than flowers. Pedicels erecto-patent, 1-6 mm. Calyx c. 1.3 cm, in fruit becoming membranous and expanding to c. 2.2 x 1.7 cm; tube c. 1 cm long; lobes each with 2 triangular acuminate teeth. Corolla 17-22 mm; tube c. ½ length of corolla, narrowly cylindrical, slightly curved. Capsule l.3x 1 .3 cm; seeds c. 5 x 4 mm. Fl. 6-8. Wet meadows, fields at forest edge, 1100-2300 m.
Described from Halle (Germany).
N. Turkey, S. & E. Anatolia. A1(E) Kirklareli: unloc, Urumov. A3 Bolu : Abant G., 1400 m, D. 37165! A4 Çankiri: Küçük Ilgaz Da., 1700-1750 m, Hııb.-Mor. 17441. A7 Trabzon: Hamsiköy. T.Baytop (ISTE 14239)! A8 Rize: d. Ikizdere, Başköy to Cermanin Y., 2300 m, D. 21033! A9 Çoruh: Ardanuç to Kordevan Da., 1400 m, D. 30156! B8 Erzurum: 29 km from Hinis to Pasinler, 1800 m, D. 46380! B9 Bitlis: Tatvan to Tuğ, 1800 m, D. 22320! B10 Van: 5 km N.E. of Başkale, 2150 m, D. 45937! C5 Adana: Komyschli (Kamişli) Su, Kotschy 1859: 231!
N., W. & C. Europe, Italy (?), Jugoslavia, Bulgaria, Caucasia, Crimea, Palestine, N. Iran, Turkestan. Although several other species of Rhinanthus have been recorded from Turkey (such as R. minor L., 'major' L., 'crista-galli' L., tournefortii Jaub. & Spach and R. glandulosus Simonkai), these all seem to be based on misidentifications. The only taxon I have recognized seems in no way to differ from the widespread European subspecies. Its nomenclature and status can only be finalized by a more broadly-based revision than presently is at hand.