Scrophulariaceae

S. cinerascens Boiss. in Tchihat., Asie Min. Bot. 2: 16 (1860). Syn: S. laciniciata sensu C. Koch in Linnaea 22:710 (1849) non Waldst. & Kit. (1803-4) var. cicleno-phora C.Koch loc. cit.; 5. micradenia Freyn in Bull. Herb. Boiss. ser. 2, 1:277 (1901)! 5. grossiheimii Schischkin in Beih. Bot. Centr. 44(2):238 (1927)! Map 46, p. 621.
Perennial; whole plant except calyx very densely glandular, often grey-green. Stems numerous, ascending, 10-50 cm. Leaves numerous, lamina narrowly ovate to oblong, margin dentate to ± deeply pinnatifid. segments often narrow. Inflorescence aphyllous; cymes 1-5-flowered. Peduncle 7-16 mm. Alar pedicels. 0.5-4.5 mm. longer or shorter than 1-2.5 mm bracteoles. Calyx lobes 1.5-2.5 x 1-2 mm, glandular to glabrous; scarious margin 0.2-0.4 mm broad, white. Corolla greenish-pink, 3.5-7 mm, upper lobes purple. Stamens included to exserted; staminode semicircular, transversely oblong, obovate-spathulate to narrowly ovate. Capsule 3.5-5 x 4-5 mm, subglobose, apiculate. dark brown to greenish-buff. Fl. 5-8. Open rocky igneous limes/one and chalk slopes and hanks. 1500-2500 m.
Type: [Turkey A7 Giresun] Ponto australi, declivitate merid. jugi Paryadres veterum (Balaban Da.) inter pagos Lisdja et Kumbetkhan, 1918-2646 m, 1858, Tchihatcheff 614 (holo. G!).
E. Anatolia. A8 Gümüşane: S. foot of Soğanli Da. N. of Bayburt, 1650 m. D. 32006! A9 Erzurum: Horasan to Pasinler (Grossheim 7: map 553). Kars: Kisir Da. above Susuz, 2000 m, D. 30527! B7 Erzincan: Sipikör nr "Pirinbahçe", Sint. 1890:3080! B8 Erzurum: c. 14 km from Erzurum to Pasinler, 2000 m, D. 47432! B9 Erzurum: c. 20 km from Horasan to Tahir, 1700 m. D. 47296! Kars: nr Iğdir (Grossheim 7: map 553). Van: nr Van. 2200 m, 12 v 1899, Kronenburg 25'a' (lectotype of 5. micradenia)!. B10 Kars: Doğubayazit to Iğdir. 2000-2500 m, Ri.x 2003!
Caucasia, Soviet Armenia. Related to S. variegata and 5. depauperate: from both of which it differs in its much more glandular habit. From S. depauperata it is additionally distinguished by the narrow calyx lobes never broader than long, and by the slightly narrower scarious margin to the calyx lobes. Forms of S. cinerascens from N. Anatolia and Caucasia tend to have the staminode semicircular to transversely oblong (more rarely ovate), and have been called 5. gross-heimii. Southern forms, including the type of S. micradenia. have smaller, broadly obovate-spathulate to narrowly ovate staminodes. Transitional forms occur and the variation in staminode morphology does not seem correlated with other characters.